• Title/Summary/Keyword: SHADE TOLERANCE

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Germination and Growth of Oaks (Quercus serrata, Q. mongolica, Q. variabilis) Seedlings by Gradient of Light Intensity and Soil Moisture (광도와 토양수분 구배(勾配)에 따른 참나무류(Quercus Serrata, Q. mongolica, Q. variabilis)치수(稚樹)의 발아 및 성장)

  • Beon Mu-Sup
    • Korean Journal of Agricultural and Forest Meteorology
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    • v.2 no.4
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    • pp.183-189
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    • 2000
  • This study was carried out to analyze ecophysiological responses for seedling of Quercus serrata, Quercus mongolica and Quercus variabilis that are the typical species of deciduous broadleaved forests in Korea. And executed experiments in the climatic control room to provide necessary information to ecological forest management and methods of natural regeneration through the analysis of seedling responses. The details of experimental analysis were growth processes of 4 months after seeding that vary with the condition of three light intensity[relative light intensity(RLI) 8%, 20%, 52%] and three soil moisture[water suction(WS) Ψ=100 hPa, Ψ=280 hPa, Ψ=330 hPa] gradient, growth factors after harvesting and the nutrition condition of leaves. The results of this study are followings: 1) Early growth was prosperous after germination for the species which have more weight of acorn. 2) The formation of lammas shoot was favourable with Q. variabilis and Q. mongolica. And the rate of the occurrence was the highest in the RLI 20%, and it was remarkably reduced in the RLI 8%. 3) As the height growth of seedling of all 3 species were greater in the RLI 20% and 8% than that of the RLI 52%, they showed strong shade tolerance. 4) The increase of light intensity promoted the diameter at root collar growth, and development of main and lateral roots with all 3 species. 5) It showed that the increase of light intensity in the experimental radiation condition raised special leaf area weight(mg/cm$^2$) and leaf area productivity(mg/cm$^2$). Consequently, these resulted in the increase of leaf thickness and total dry biomass per the unit area of leaf. 6) As the increase of light intensity, the minerals contents of leaves such as N, P and K were lowered, and the increase of soil moisture resulted in the increase of P, K, Ca and Mg.

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A Study on the Visible Injury of some Herbaceous Plants by $SO_2$ gas (수종(數種) 초본류(草本類)의 $SO_2$ 가스에 의한 가시피해특징(可視被害特徵)에 관(關)한 연구(硏究))

  • Kim, Jeong-Gyu;Lim, Soo-Kil;Kim, Jae-Bong
    • Korean Journal of Environmental Agriculture
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    • v.7 no.1
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    • pp.43-51
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    • 1988
  • $SO_2$ gas injuries for 19 varieties of 10 species of plants were investigated within a environmentally controlled growth chamber. Visible injuries were observed when exposing the plants either 0, 0.2, 0.4, 0.7 or 1.5ppm of $SO_2$ gas for 5 days from 9 : 00 to 17 : 00 everyday. The results obtained were as follows: 1. Light brown spots were shown on Raphanus sativus(Chunchu, Taeback, Jinjudaepyung), Brassica campestris(Manchun, Josaengmiho, Seoul, Jeonseung), and Capsicum annum(Searona, Hongsanho, Hongilpum); brown spots on Sesamum indicum(Pungnyun, Kwangsan); reddish brown shades on Cleosia cristata; and black brown spots on Perilla frutescens, Calendula officinalis, Chrysanthemum morifolium, and Salvia splendens, respectively. 2. The spotty injuries induced by $SO_2$ gas injuries for 19 varieties of 10 species of plants were investigated within a environmentally controlled growth chamber. Visible injuries were observed when exposing the plants either 0, 0.2, 0.4, 0.7 or 1.5ppm of $SO_2$ gas for 5 days from 9 : 00 to 17 : 00 everyday. The results obtained were as follows: 1. Light brown spots were shown on Raphanus sativus(Chunchu, Taeback, Jinjudaepyung), Brassica campestris(Manchun, Josaengmiho, Seoul, Jeonseung), and Capsicum annum(Searona, Hongsanho, Hongilpum); brown spots on Sesamum indicum(Pungnyun, Kwangsan); reddish brown shades on Cleosia cristata; and black brown spots on Perilla frutescens, Calendula officinalis, Chrysanthemum morifolium, and Salvia splendens, respectively. 2. The spotty injuries induced by SO₂ gas on Raphanus sativus, Callistephus chinensis, Capsicum annum, Perilla frutescens, Calendula officinalis, Salvia splendens, and Sesamum indicum; the many small spots on Chrysanthemum morifolium; and the brown shade on Celosia cristata appeared on the upper surface of the intervein, on the leaf apex area and on the entire upper surface of leaves, respectively. 3. Visual injuries of Capsicum annum(Chunchu), Perilla frutescens, Sesamum indicum(Pungnyun, Kwangsan), and Salvia splendens were developed at 0.4ppm of SO₂ gas fumigation. Brassica campestris(Jeonseung), Capsicum annum(Searona, Hongilpum), and Cleosia cristata, however, showed the injury at 1.5ppm of $SO_2$ gas concentration. 4. Based on the tolerance grade(time when the 1st injury appeared), Raphanus sativus, Perilla frutescens, Sesamum indicum, and Salvia splendens were the most sensitive plants and Chrysanthemum morifolium, Callistephus chinensis, Cleosia cristata, and Calendula officinalis were the plants most tolerant of $SO_2$ gas. gas on Raphanus sativus, Callistephus chinensis, Capsicum annum, Perilla frutescens, Calendula officinalis, Salvia splendens, and Sesamum indicum; the many small spots on Chrysanthemum morifolium; and the brown shade on Celosia cristata appeared on the upper surface of the intervein, on the leaf apex area and on the entire upper surface of leaves, respectively. 3. Visual injuries of Capsicum annum(Chunchu), Perilla frutescens, Sesamum indicum(Pungnyun, Kwangsan), and Salvia splendens were developed at 0.4ppm of $SO_2$ gas injuries for 19 varieties of 10 species of plants were investigated within a environmentally controlled growth chamber. Visible injuries were observed when exposing the plants either 0, 0.2, 0.4, 0.7 or 1.5ppm of $SO_2$ gas for 5 days from 9 : 00 to 17 : 00 everyday. The results obtained were as follows: 1. Light brown spots were shown on Raphanus sativus(Chunchu, Taeback, Jinjudaepyung), Brassica campestris(Manchun, Josaengmiho, Seoul, Jeonseung), and Capsicum annum(Searona, Hongsanho, Hongilpum); brown spots on Sesamum indicum(Pungnyun, Kwangsan); reddish brown shades on Cleosia cristata; and black brown spots on Perilla frutescens, Calendula officinalis, Chrysanthemum morifolium, and Salvia splendens, respectively. 2. The spotty injuries induced by SO₂ gas on Raphanus sativus, Callistephus chinensis, Capsicum annum, Perilla frutescens, Calendula officinalis, Salvia splendens, and Sesamum indicum; the many small spots on Chrysanthemum morifolium; and the brown shade on Celosia cristata appeared on the upper surface of the intervein, on the leaf apex area and on the entire upper surface of leaves, respectively. 3. Visual injuries of Capsicum annum(Chunchu), Perilla frutescens, Sesamum indicum(Pungnyun, Kwangsan), and Salvia splendens were developed at 0.4ppm of SO₂ gas fumigation. Brassica campestris(Jeonseung), Capsicum annum(Searona, Hongilpum), and Cleosia cristata, however, showed the injury at 1.5ppm of $SO_2$ gas concentration. 4. Based on the tolerance grade(time when the 1st injury appeared), Raphanus sativus, Perilla frutescens, Sesamum indicum, and Salvia splendens were the most sensitive plants and Chrysanthemum morifolium, Callistephus chinensis, Cleosia cristata, and Calendula officinalis were the plants most tolerant of $SO_2$ gas. gas fumigation. Brassica campestris(Jeonseung), Capsicum annum(Searona, Hongilpum), and Cleosia cristata, however, showed the injury at 1.5ppm of $SO_2$ gas concentration. 4. Based on the tolerance grade(time when the 1st injury appeared), Raphanus sativus, Perilla frutescens, Sesamum indicum, and Salvia splendens were the most sensitive plants and Chrysanthemum morifolium, Callistephus chinensis, Cleosia cristata, and Calendula officinalis were the plants most tolerant of $SO_2$ gas.

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Studies on the Shade Tolerance, Light Requirement, and Water Relations of Economic Tree Species(III) - Analysis of Pressure-Volume Curves on the Changes of Tissue Water Relations of Five Deciduous Hardwood Species Subjected to Artificial Shading Treatments - (주요경제수종(主要經濟樹種)의 내음성(耐陰性) 및 광선요구도(光線要求度)와 수분특성(水分特性)에 관한 연구(III) - 인공피음처리하(人工被陰處理下)에서 자라는 활엽수(闊葉樹) 5수종(樹種)의 수분특성(水分特性) 변화(變化)에 대한 P-V곡선(曲線) 분석(分析) -)

  • Choi, Jeong Ho;Kwon, Ki Won
    • Journal of Korean Society of Forest Science
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    • v.90 no.4
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    • pp.524-534
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    • 2001
  • The pressure-volume curve parameters were investigated to elucidate the effects of shading treatment on the water relations of the one year old seedlings of Betula platyphylla var. japonica, Betula schmidtii, Zelkova serrata, Acer mono and Prunes sargentii subjected to five levels of artificial shading treatments. The osmotic potentials at full turgor(${\phi}_{{\pi}o}$) measured under full sunlight changed with species and growing season in the ranges of -1.04~-1.27MPa, -1.03~-1.48MPa, -0.94~-1.44MPa in first year treatment, and -0.90~-1.37MPa, -1.05~-1.79MPa, -0.99~-1.30MPa in second year treatment in June, July, and September, respectively. The osmotic potentials at full turgor increased with increment of shading level in the ranges of -0.90~-1.79MPa in full sunlight and -0.58~-1.23MPa in nearly full shading level(E) through the growing seasons in all the species studied. The osmotic potentials at turgor loss point(${\phi}_{{\pi}p}$) measured in full sunlight changed in the ranges of -1.64~-2.11MPa, -1.67~-2.15MPa, -1.47~-2.11MPa, and -1.45~-2.04MPa, -1.30~-2.00MPa, -1.28~-2.33MPa in June, July, and September of first and second years, respectively. Most of ${\phi}_{{\pi}p}$ measurements were lower within about 0.5MPa in comparison with those of ${\phi}_{{\pi}o}$. The measurements of ${\phi}_{{\pi}p}$ also increased with increment of shading level, and the differences in ${\phi}_{{\pi}p}$ among shading levels were generally greater than those in ${\phi}_{{\pi}o}$ by species and by growing season. Most of the osmotic potentials at turgor loss point as like as at full turgor were lowered in July than in June and September. The measurements of relative water content at turgor lass point(RWCp) in full sunlight were in the similar ranges of 81~88%, 71~86%, 75~84%, and 82~87, 72~84%, 76~86% in June, July, and September of first and second years, respectively. The RWCp were a little higher in A. mono and P. sargentii than in B. platyphylla var. japonica, B. schmidtii, and Z. serrata. The RWCp also decreased from 71~88% in full sunlight to 48~77% in nearly full shading treatment with increment of shading level. Even if there were some exceptions by species or by growing season, the shading effects on the changes in some P-V parameters were distinctly observed in the present study. The change in P-V parameters following shading treatment may be presumably inferred on the changes in solute accumulation, membrane elasticity, symplasmic water volume, and so on. But much more experiments should be necessarily continued for getting detailed informations on the physiological mechanism of shading effects relating to the changes in P-V parameters.

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The Study on the Height Characteristics of Abies Nephrolepis Community in South Korea - In the Case of Seorak·Odae·Taebaek National Park - (우리나라 분비나무의 수고 특성 연구 - 설악·오대·태백산국립공원을 대상으로 -)

  • Jin-Won Kim;Ho-Young Lee;Young-Moon Chun;Choong-Hyeon Oh
    • Korean Journal of Environment and Ecology
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    • v.38 no.2
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    • pp.169-177
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    • 2024
  • This study investigated whether population dynamic analysis based on the height characteristics of Abies nephrolepis was feasible. It was necessary because existing population dynamic analyses based on age and diameter at breast height (DBH) made it difficult to reflect the slow growth characteristics of Abies nephrolepis in harsh environments of high altitudes. The limitations of population dynamics analysis based on the age and DBH distribution of Abies nephrolepis in Seoraksan, Odaesan, and Taebaeksan National Parks, where Abies nephrolepis populations are representative, were verified, and the characteristics of height growth were investigated to comprehensively analyze whether a vertical structure based on height could reveal the population dynamics. The result of this study showed some limitations in understanding the population dynamics of Abies nephrolepis based on age distribution due to practical difficulties in sampling all trees and variations in age distribution within the same community depending on factors such as light conditions. Moreover, it was challenging to differentiate the distribution of DBH classes at fine levels, making it difficult to reflect the rapid growth characteristics of Abies nephrolepis when light conditions become suitable after prolonged stays in smaller DBH classes under shade conditions. However, a comprehensive analysis of the height characteristics of Abies nephrolepis revealed that the density corresponding to the population dynamic characteristics of Abies was high and adequately reflected the predominant tree death at similar height stages, as well as the U-shaped population dynamics at the lower stratum. Moreover, it was possible to identify a transition point in height values under shaded conditions, where the annual growth of Abies nephrolepis individuals in the lower stratum increases significantly, indicating that Abies nephrolepis individuals can escape from competition with other shrubs and undergo vigorous growth only at this height level. Therefore, this study confirmed that a vertical structure based on height can be utilized to understand the population dynamics of Abies nephrolepis in high altitudes, and it is expected that future studies on height characteristics can intuitively reveal the maintenance status of Abies nephrolepis populations in the field.

Water Physiology of Panax ginseng. 1. Habitat observation. cultural experience, weather factors and characteristics of root and leaf (인삼의 수분생리 1. 자생지관찰.재배 경험.기상요인과 근 및 엽의 특성)

  • Park, Hoon
    • Journal of Ginseng Research
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    • v.4 no.2
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    • pp.197-221
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    • 1980
  • Habitat observation, cultural experience of old and present plantation, weather factors in relation to crop stand and water physiology of root and leaf were reviewed. According to habitat observation ginseng plants love water but plate wit talus well grow at drained place with high moisture content in air and soil while ginseng plants were not found in dry or wet place. According to cultivation experience ginseng plants require abundant water in nursery and main field but most old planters believe that ginseng plaints are draught-loving thus require little water. The experience that rain especially in summer i.e unfavorable might be due to mechanical damage of leaves arid leaf disease infection, or severe leaf fall which is caused by high air temperature and coinsided with rain. According to crop stand observation in relation to weather factors abunsant water increased each root weight but decreased total yield indicating tile increase of missing root rate. Rain in summer was unfavorable too. Though rain in June was favorable for high yield general experience that cloudy day and rain were unfavorable might be due to low light intensity under shade. Present leading planters also do loot consider the importance of water in main field. Water content is higher in top than in root and highest in central portion of root and in stem of top. For seedling the heavier the weight of root is tile higher the water content while it reveries from two years old. Water potential of intact root appeared to be -2.89 bar suggesting high sensitivity to water environment. Under water stress water content severly decreased only in leaf. Water content of leaf appeared to be 78% for optimum, below 72% for functional damage and 68% for perm anent wilting. Transpiration or curs Principally through stomata in lower side of leaf thus contribution of upper side transpiration decreased with the increase of intensity. Transpiration is greater in the leaves grown under high light intensity. Thus water content is lower with high light inte nsity under field condition indicating that light is probable cause of water stress in field. Transpiration reached maximum at 10K1ut The decrease of transpiration at higher temperature seems to be due to the decrease of stomata aperture caused by water stress. Severe decrease of photosynthesis under water stress seems to be principally due to functional damage which is not caused by high temperature and Partly due to poor CO2 supply. Water potential of leaf appeared to be -16.8 bar suggesting weakness in draught tolerance. Ginseng leaves absorb water under high humidity. Water free space of leaf disc is %mailer than that of soybean leaf and water uptake appears to be more than two steps.

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Growth Difference between the Seedlings of Quercus serrata and Q. aliena under light, moisture and nutrient Gradients (광, 수분, 영양소에 따른 졸참나무와 갈참나무 유식물의 생육 차이)

  • Lim, Hoon;Kim, Hae-Ran;You, Young-Han
    • Journal of Wetlands Research
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    • v.14 no.2
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    • pp.237-242
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    • 2012
  • In order to determine the ecological characteristics of Quercus serrata and Q. aliena, which are potential natural vegetation of riverine in Korea, we cultivated the seedlings of two oak species under light, soil moisture and nutrient gradients from April to October in glasshouse. Then, we measured aboveground, belowground and total plant biomass and analyzed the differences in growth between two oak species. The two oak species showed decreasing growth with lower light intensity, but reduction in growth of Q. aliena was greater than that of Q. serrata. Q. serrata and Q. aliena had a constant growth state under soil moisture gradients, but Q. aliena grew well more than Q. serrata in lower soil moisture gradient. As soil nutrient availability decreased, the growth of Q. aliena was greater than Q. serrata. These results mean that the growth of two oak species is not affected by soil moisture, but the part of growth in Q. serrata may be positively affected in lower soil moisture condition. Also, Q. serrata has strong shade tolerance and the ability to adapt high nutrient condition relative to Q. aliena. Thus, Q. serrata and Q. aliena will be advantageous for the growth in stream ecosystem due to high light availability, but Q. serrata will be distributed near the lowlands by the water environment. Also, because the two oak species need high soil nutrient, it is important to preserve herbaceous vegetation to prevent nutrients that flowed in the streams.

Growth and Physiological Response in Container of Daphniphyllum macropodum by Shading Level (피음 수준에 따른 굴거리나무의 용기 내 생장 및 생리적 반응)

  • Song, Ki Seon;Kim, Jong Jin
    • Journal of Korean Society of Forest Science
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    • v.106 no.1
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    • pp.33-39
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    • 2017
  • This study was carried out in order to closely examine the growth and the physiological response by shading levels of a container seedling of Daphniphyllum macropodum which is known as the species of having shade tolerance in a warm temperature evergreen broad-leaved tree. The shading treatment was regulated with the shading level of full sunlight, and 35%, 55%, 75% of full sunlight. As a result of surveying the height and the root collar diameter growth by the shading level, the height was the highest with 13.2 cm under 35% of shading. The root collar diameter was the highest with 4.99 mm under full sunlight. The organ-based (leaves, shoot and root) dry mass production and the whole dry mass production were higher with 1.38 g, 0.47 g, 0.87 g, 2.72 g, respectively, in order of 35% shading. As for chlorophyll a, b, and the total chlorophyll content, the highest chlorophyll content was shown under 75% of shading with the relatively highest shading level. The ratio of chlorophyll a to chlorophyll b (a/b) was the highest under 35% of shading. The photosynthetic rate, stomatal conductance and water use efficiency were the highest with $12.3{\mu}molCO_2{\cdot}m^{-2}{\cdot}s^{-1}$, $0.27molH_2O{\cdot}m^{-2}{\cdot}s^{-1}$, $3.53{\mu}molCO_2{\cdot}mmol^{-1}H_2O$, respectively, under 35% of shading. As a result of surveying the whole experiment, D. macropodum grows nicely by maintaining 35% shading.

Proper Light Intensity, Potting Media and Fertilization Level for Potted Hepatica asiatica Nakai (노루귀의 분화재배를 위한 적정 광도, 분용토 및 시비수준)

  • Jeong, Kyeong Jin;Jeon, Hyeon Sik;Chon, Young Shin;Yun, Jae Gill
    • Horticultural Science & Technology
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    • v.33 no.1
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    • pp.24-30
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    • 2015
  • This study was conducted to select proper light intensity, pot media, and fertilization level for potted Hepatica asiatica Nakai native to Korea. The plants were grown under various light intensities (shading rate, 52, 82, 90, 97%) imposed by shading net. Plants grown with 52% shading showed a low survival rate (65%). Survival rate increased as shading increased, with over 80% survival in shading above 90%. Growth indexes such as fresh weight and leaf number did not show any significant difference between shading treatments. Plants grown in a soil mixture of decomposed granite:fertilizer-amended media:Kanumatsuchi (60:10:30, v/v/v) or river sand:fertilizer-amended media:bark (50:20:30) showed over 85% survival. However, plants grown in a soil mixture of river sand:fertilizer-amended media:Kanumatsuchi (50:30:20) or upland:river sand (40:60) showed very low survival, below 60%. Leaf number and plant height were the highest in a soil mixture of decomposed granite:fertilizer-amended media:Kanumatsuchi (60:10:30) as well. To select a proper fertilization level for H. asiatica, hyponex solution diluted 1,000- or 2,000-fold were applied weekly or biweekly. The survival rate was lowest at weekly application with 1,000-fold diluted solution, and no significant difference was observed between other treatments. In conclusion, H. asiatica exhibits preferences for very low light intensity and soil with air permeability, and is adaptable to a broad range of fertilization levels.

Vegetation structure and distribution characteristics of Symplocos prunifolia, a rare evergreen broad-leaved tree in Korea

  • Kim, Yangji;Song, Kukman;Yim, Eunyoung;Seo, Yeonok;Choi, Hyungsoon;Choi, Byoungki
    • Journal of Ecology and Environment
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    • v.44 no.4
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    • pp.275-285
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    • 2020
  • Background: In Korea, Symplocos prunifolia Siebold. & Zucc. is only found on Jeju Island. Conservation of the species is difficult because little is known about its distribution and natural habitat. The lack of research and survey data on the characteristics of native vegetation and distribution of this species means that there is insufficient information to guide the management and conservation of this species and related vegetation. Therefore, this study aims to identify the distribution and vegetation associated with S. prunifolia. Results: As a result of field investigations, it was confirmed that the native S. prunifolia communities were distributed in 4 areas located on the southern side of Mt. Halla and within the evergreen broad-leaved forest zones. Furthermore, these evergreen broad-leaved forest zones are themselves located in the warm temperate zone which are distributed along the valley sides at elevations between 318 and 461 m. S. prunifolia was only found on the south side of Mt. Halla, and mainly on south-facing slopes; however, small communities were found to be growing on northwest-facing slopes. It has been confirmed that S. prunifolia trees are rare but an important constituent species in the evergreen broad-leaved forest of Jeju. The mean importance percentage of S. prunifolia community was 48.84 for Castanopsis sieboldii, 17.79 for Quercus acuta, and 12.12 for Pinus thunbergii; S. prunifolia was the ninth most important species (2.6). Conclusions: S. prunifolia can be found growing along the natural streams of Jeju, where there is little anthropogenic influence and where the streams have caused soil disturbance through natural processes of erosion and deposition of sediments. Currently, the native area of S. prunifolia is about 3300 ㎡, which contains a confirmed population of 180 individual plants. As a result of these low population sizes, it places it in the category of an extremely endangered plant in Korea. In some native sites, the canopy of evergreen broad-leaved forest formed, but the frequency and coverage of species were not high. Negative factors that contributed to the low distribution of this species were factors such as lacking in shade tolerance, low fruiting rates, small native areas, and special habitats as well as requiring adequate stream disturbance. Presently, due to changes in climate, it is unclear whether this species will see an increase in its population and habitat area or whether it will remain as an endangered species within Korea. What is clear, however, is that the preservation of the present native habitats and population is extremely important if the population is to be maintained and expanded. It is also meaningful in terms of the stable conservation of biodiversity in Korea. Therefore, based on the results of this study, it is judged that a systematic evaluation for the preservation and conservation of the habitat and vegetation management method of S. prunifolia should be conducted.

Characteristics of Seedling Quality of Daphniphyllum macropodum 2-year-old Container Seedlings by Shading Level (굴거리나무 2년생 용기묘의 피음수준별 묘목품질 특성)

  • Song, Ki Seon;Choi, Kyu Seong;Sung, Hwan In;Jeon, Kwon Seok;An, Kyoung Jin;Kim, Jong Jin
    • Journal of Korean Society of Forest Science
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    • v.104 no.3
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    • pp.390-396
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    • 2015
  • This study was carried out in order to closely examine quality index by shading level of 2-year-old (1-1 seedling) container seedling of Daphniphyllum macropodum which is known as the species of having shade tolerance that is evergreen broad leaved tree in the warm temperate region. The shading treatment was regulated with the shading level of full sunlight, and 35%, 55%, 75%, 95% of full sunlight. As a result of surveying growth according to the shading level, both height and root collar diameter were surveyed to be the highest with 45.1 cm and 8.22 mm, respectively, under 75% of shading. The next was surveyed to be 43.2 cm & 8.05 mm and 42.5 cm & 7.98 mm, respectively, in order of 35% and 55% in shading. Leaf, shoot, root, and whole dry mass production were the highest under 75% of shading. The next was higher in leaf, stem, and whole dry mass production under 55% of shading. A root was higher under 35% of shading in the next. H/D ratio was the range of 5.29~5.35 under the 35~75% shading that showed the relatively high height and root collar diameter. T/R ratio was the lowest with 1.17 under 35% of shading. It was 0.41 under 95% of shading as for LWR, 0.24 under 75%-95% of shading as for SWR, and 0.46 under full sun and 35% shading as for RWR. QI was the highest with 3.74 under 75% of shading. As a result of surveying the whole experiment, it is concluded that the production of D. macropodum seedling is more effective under 75% shading