• Title/Summary/Keyword: etten

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Two Specificity markers etten and han in Korean (두 특정성 표지 '어떤'과 '한'에 대한 고찰)

  • Yeom, Jae-Il
    • Language and Information
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    • v.11 no.2
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    • pp.23-47
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    • 2007
  • In this paper I discuss two markers for specificity, etten and han, in Korean. First, I discuss their structural properties and claim that etten is a determiner, and han is a numeral preceded by an implicit existential quantifier. Then I discuss four uses of etten and two uses of han, and show when they are used as specificity markers. There are various properties of the two specificity markers, some of which they have in common. I take them as properties of specificity markers in general. The properties only one of them has are taken to come from the original meanings of the two morphemes. I claim that specific indefinites range over individual concepts, which lead to the properties they have in common.

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Comparing Plant Species Diversity of Mountainous Deserts - Successes and Pitfalls

  • Van Etten, Eddie J.B.
    • The Korean Journal of Ecology
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    • v.27 no.2
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    • pp.79-86
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    • 2004
  • An extensive study of the vegetation characteristics of the Hamersley Ranges, a mountainous desert area of north-west Australia, facilitated the comparison of plant species diversity measures with mountainous deserts of other parts of the world. Alpha diversity was defined as the number of species co-existing at local scales and was found to average 18 species per 0.1 ha for the Hamersley Ranges. This was found to be similar to seven other mountainous deserts in North and South America, and southern Africa. Variation in alpha diversity between these deserts was found to considerably lower than within deserts, suggesting that local processes control species richness at local scales. Beta diversity, defined here as turnover in species composition at various spatial scales, can be measured in many ways. For the Hamersley Ranges, Wilson's β ranged from 1.2 to 1.6 for five sites along a topographic gradient, whereas Whittaker's β between different plant communities was found to average 0.93. Comparable data was not found for other desert areas, but comparisons to non-desert areas suggest beta diversity within landscapes is relatively high and is likely to reflect the considerable landform heterogeneity of the Hamersley Ranges. 55∼70% of species were shared between different landscapes of the Hamersley Ranges; comparisons to other regions suggest beta diversity at this scale is relatively low. Gamma diversity, the number of species over large spatial extents, was successfully compared using regression analysis of the log-log species - area relationship. This revealed that the northern Sonoran desert has significantly less species than the Nama (inland) Karoo and Hamersley Ranges over medium spatial extents, but species numbers were similar at a regional scale. Several constraints to the valid comparison of species diversity were identified, including lack of standardisation of sampling techniques, the wide range of measures employed, general lack of published data, and the influence of the various components of spatial scale on most diversity measures. Recommendations on how to improve future comparative work are provided.

Functional Implication of the tRNA Genes Encoded in the Chlorella Virus PBCV-l Genome

  • Lee, Da-Young;Graves, Michael V.;Van Etten, James L.;Choi, Tae-Jin
    • The Plant Pathology Journal
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    • v.21 no.4
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    • pp.334-342
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    • 2005
  • The prototype Chlorella virus PBCV-l encodes 11 tRNA genes and over 350 protein-encoding genes in its 330 kbp genome. Initial attempts to overexpress the recombinant A189/192R protein, a putative virus attachment protein, in E. coli strain BL21(DE3) SI were unsuccessful, and multiple protein bands were detected on Western blots. However, the full-length A189/192R recombinant protein or fragments derived from it were detected when they were expressed in E. coli BL21 CodonPlus (DE3) RIL, which contains extra tRNAs. Codon usage analysis of the a189/192r gene showed highly biased usage of the AGA and AVA codons compared to genes encoded by E. coli and Chlorella. In addition, there were biases of XXA/U($56\%$) and XXG/ C($44\%$) in the codons recognized by the viral tRNAs, which correspond to the codon usage bias in the PBCV-1 genome of XXA/U ($63\%$) over those ending in XXC/G ($37\%$). Analysis of the codon usage in the major capsid protein and DNA polymerase showed preferential usage of codons that can be recognized by the viral tRNAs. The Asn (AAC) and Lys (AAG) codons whose corresponding tRNA genes are duplicated in the tRNA gene cluster were the most abundant (i.e., preferred) codons in these two proteins. The tRNA genes encoded in the PBCV-l genome seem to play a very important role during the synthesis of viral proteins through supplementing the tRNAs that are frequently used in viral proteins, but are rare in the host cells. In addition, these tRNAs would help the virus to adapt to a wide range of hosts by providing tRNAs that are rare in the host cells.