• Title/Summary/Keyword: Taxus cuspidata

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The Analysis of Vegetation-Environment Relationships of the Coniferous Forests in Subalpine Districts of Mt. Chiri by TWINSPAN and CCA (TWINSPAN 및 CCA에 의한 지리산(智異山) 아고산대(亞高山帶) 침엽수림군집(針葉樹林群集)과 환경(環境)의 상관관계(相關關係) 분석(分析))

  • Lee, Soo Won;Lee, Kang Young;Song, Ho Kyung
    • Journal of Korean Society of Forest Science
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    • v.86 no.3
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    • pp.279-287
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    • 1997
  • This study was carried out to identify the analysis of vegetation-environment relationships of the coniferous forests in subalpine districts of Mt. Chiri by two-way indicaton species analysis(TWINSPAN) and CCA. The results are summarized as follows; The subalpine coniferous forest in Mt. Chin was classified Abies koreana community group by the TWINSPAN method. The Abies koreana community group was classified Abies koreana-Pinus densiflora, Abies koreana-Quercus mongolicd and Abies koreana-Picea jezoensis. The Abies koreana-Quercus mongolica community classified into two subcommunities, typical and Pinus koraiensis subcommunity. The Abies koreana-Picea jezoensis community classified into three subcommunities such as typical, Taxus cuspidate and Cornus controversa subcommunity. The subalpine conifers commnities in Mt. Chiri could be classified into six vegetation units, which consisted of three vegetation communities and five subcommunities. The relationship between vegetation community and influencing environmental factors was analyzed by using the CCA ordination method. It was found that altitude, concentrations of $Ca^{++}$ and $Mg^{++}$ in soil were, major environmental variables, which influence the distribution of vegetation community in the first axis, and $Ca^{++}$ and altitude in the second axis. In the relations of communities and environmental factors by CCA, the optimal ecological habitats of Abies koreana-Pinus densiflora community could be located at low elevations, in which $Ca^{++}$ and $Mg^{++}$ are rich. Abies koreana-Quercus mongolica community prefers the sites in mid-elevation zone, in which $Ca^{++}$ and $Mg^{++}$ are medium level in soil. The optimum site for Abies koreana-Pinus koraiensis community and Abies koreana-Taxus cuspidata community could be high elevations in which $Ca^{++}$ and $Mg^{++}$ are poor. Abies koreana-Cornus controversa community is found in the site at low elevations where $Ca^{++}$ and $Mg^{++}$ are medium.

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Study on the DBH Analysis and Forest Succession of Pinus densiflora and Quercus mongolica Forests (소나무림(林)과 신갈나무림(林)의 흉고직경급(胸高直徑級) 분석(分析)과 천이(遷移)에 관(關)한 연구(硏究))

  • Song, Ho Kyung;Jang, Kyu Kwan
    • Journal of Korean Society of Forest Science
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    • v.86 no.2
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    • pp.223-232
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    • 1997
  • One hundred Sixty two quadrats were sampled from natural forests, in Kangwon-do and Kyungsangbuk-do. On the basis of Mueller-Dombois & Ellenberg method, field survey, which accounts for the characters of the species structure of Pinus densiflora community and Quercus mongolica community, was carried out between 1991 and 1994. To analyze the data, CCA ordination and D.B.H, analysis were used. 1. DBH analysis showed that Pinus densiflora community is likely replacing Quercus mongolica or Quercus variabilis community. DBH analysis also showed that Quercus mongolica - Lindera obtusiloba community balances among the Quercus mongolica, Fraxinus rhynchophylla, and Tilia amurensid in the canopy, and that Quercus mongolica-Abies nephrolepis community, mixed forest balances among the Quercus mongolica, Pinus koraiensis, and Abies nephrolepis in the canopy. In general, Quercus mongolica seems to remain as a dominant species. 2. A certain correlation seems to exist between nutrients types and Pinus densiflora community. Quercus mongolica prefered to grow in the medium elevation area that has medium level of total nitrogen, organic matter, and C.E.C. Pinus densiflora, Quercus serrata and Quercus variabilis were found in the low elevation area that has low level of total nitrogen, organic matter, and C.E.C. 3. There seemed to be a certain correlation between nutrients types and Quercus mongolica community. Quercus mongolica and Acer pseudosieboldianum was distributed in the slope area that has medium level of $Mg^{{+}{+}}$, $Ca^{{+}{+}}$, and C.E.C., while Abies nephrolepis, Taxus cuspidata and Pinus koraiensis in the high elevation and ridge area that has low level of $Mg^{{+}{+}}$, $Ca^{{+}{+}}$, and C.E.C. Carpinus laxiflora and Abies holophylla were found in the medium elevation area that has medium level of $Mg^{{+}{+}}$, $Ca^{{+}{+}}$, and C.E.C., while Kalopanax pious, Carpinus cordata and Acer truncatum in the moderately moist and valley area that has high level of $Mg^{{+}{+}}$, $Ca^{{+}{+}}$, and C.E.C. Pinus densiflora and Fraxinus rhynchophylla seemed to prefer the slightly dry, low elevation and ridge area.

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Characteristics of Breeding Birds Community in Relation to the Forest Environment in Deogyusan National Park (덕유산국립공원의 산림환경에 따른 번식기 조류군집 특성)

  • Yu, Jae-Pyoung;Jin, Seon-Deok;Kim, Hyoun-Sook;Lee, Joon-Woo;Paek, Woon-Kee;Song, Ho-Kyung
    • Korean Journal of Environment and Ecology
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    • v.25 no.2
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    • pp.132-144
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    • 2011
  • We studied the forest environment and characteristics of bird community between April and September of 2010 on the Hyangjeokbong area(A), Baekryunsa area(B) and Anseong area(C), all in the Deogyusan National Park. Hyangjeokbong area of the high latitude ridge was characterized by the Taxus cuspidata and Abies koreana etc., and deciduous broadleaf forest species, such as the Quercus mongolica, with abundance of subalpine zone coniferous trees, the Hemerocallis fulva and the Rhododendron schlippenbachii, while the areas Baekryunsa and Anseong were primarily characterized by deciduous broadleaf forests, with the valleys showing Fraxinus mandshurica colonies. In terms of the DBH(diameter at breast height) of trees, between 11 and 20cm showed the highest frequencies, while over 30cm was the highest in Baekryunsa area and the lowest in Hyangjeokbong area. Furthermore, in terms of coverage in relation to layers, in Hyangjeokbong area, the coverage volume of the lower layer was very high, in Baekryunsa area, the coverage volume of the middle layer between 8 and 12m was the highest, and in Anseong area, the coverage volume of the upper layer over 18m was the highest. A total of 53 bird species were observed during the study period at the Deogyusan National Park. The numbers of species and density of areas were 25 species and 45.20 Ind./km for Hyangjeokbong area, 50 species and 58.63 Ind./km for Baekryunsa area and 35 species and 66.89 Ind./km for Anseong area, with Baekryunsa area showing the highest number of species and Anseong area showing the highest level of density. In terms of dominant species, in Hyangjeokbong area, Cettia diphone, along with species which inhabit in grassland and shrubs, were the dominant species, and species which live in the canopy layer, including the Aegithalos caudatus, were found to be dominant in Baekryunsa and Anseong areas. For guild structure, bush nesting guild and bush foraging guild species were the highest in all areas, and the hole nesting guild and the air foraging guild species showed the lowest proportion. This seems to be the result of the low number of trees with diameter at breast height of over 30cm, which results in the lack of nesting grounds for hole nesting guild species.

Potent Anticarcinogenic Action of Moutan radix for Mouse Ascites Cancer Induced by Mouse Sarcoma 180 Cells (Moutan radix의 mouse sarcoma 180 cell로 유발한 mouse ascites cancer에 대한 항암효과)

  • Bahn, Kyeong-N.;Lee, Eun-J.;Yang, Min-S.;Kim, Jeong-O.;Ha, Yeong-L.
    • Applied Biological Chemistry
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    • v.38 no.4
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    • pp.364-369
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    • 1995
  • Anticarcinogenic activity of Moutan radix for mouse ascites cancer induced by mouse Sarcoma 180 (S-180) cells was investigated. Methanol extract of Moutan radix including other folk medicinal plants (Taxus cuspidata, Curcuma longa, Artemisia capillaris, Ligrstri fructus, and Liriope platyphylla) used to remedy or cure many chronic human diseases like cancer was fractionated into hexane, chloroform ($CHCl_3$), ethylacetate (EtOAc), and butanol (BuOH) fractions. Anticarcinogenic activity of the fractions, exhibited a strong cytotoxicity for L1210 and S-180 cells, was examined for mouse ascites cancer induced by S-180 cells. Male ICR mice (7 mice/treatment, $5{\sim}6$ weeks of age, $23{\pm}1\;g$ were injected i.p. with S-180 cells ($1{\times}10^{7}\;cell/1\;ml$ PBS). One day later, each mouse was given 0.1 ml of 10% DMSO containing sample ($30\;{\mu}g/g$ body weight) every day for 10 consecutive days. Control mice were only given 0.1ml S-180 cells and 0.1 ml 10% DMSO. Mice treated with EtOAc fraction of Moutan radix showed 28.7 days of life, which is 167% of control mice's life. Based on the dose-dependant experiment mice treated with $30\;{\mu}g$ showed longer life relative to mice treated with ootherr doses (5, 15, $60\;{\mu}g$), and mice treated with $60\;{\mu}g$ exhibited toxic symptoms. Body weight of mice treated with Moutan radix was significantly reduced relative to that of control mice (p<0.05). GC-MS analysis in conjunction with silica-gel column chromatography revealed that the EtOAc fraction contained 2-methoxylphenol, benzoic acid, 1-(4-hydroxy-3-methoxyphenyl)ethanone, 8-methyl-2,4(1H,3H)pteridinedione and 2,5-furan-dicarboxylic dimethyl ester as regards to the anticarcinogenic property of the EtOAc fraction. These results suggest that Moutan radix might be included as an anticarcinogenic medicinal plant for treatment of ascites cancer.

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A Study on the Distribution and Dynamics of Relict Forest Trees and Structural Characteristics of Forest Stands in Gangwon Province, Korea (강원지역 산림유존목의 분포, 동태 및 생육임분의 구성적 특성)

  • Shin, Joon-Hwan;Lee, Cheol-Ho;Bae, Kwan-Ho;Cho, Yong-Chan;Kim, Jun-Soo;Cho, Jun-Hee;Cho, Hyun-Je
    • Korean Journal of Environment and Ecology
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    • v.32 no.2
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    • pp.165-175
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    • 2018
  • The purpose of this study is to provide the basic data such as distribution status, growth characteristics, and the structural characteristics of forest stands for the systematic conservation and management of relict forest trees (stem girth of 300cm or larger) established naturally in Gangwon Province, Korea. The survey showed that 434 individuals of 19 species (conifers: 228 individuals of 4 species, broad-leaved trees: 206 individuals of 15 species) were distributed in Gangwon Province, and Taxus cuspidata was the most abundant among them with 203 individuals or about 46.7 % of the total. The stem girth was average of 404cm (conifers: 373cm, broad-leaves: 421cm), and Tilia amurensis with multi-stemmed growing on Sorak mountain range had the largest stem girth at 1,113cm. The average height and the crown width of relict forest trees were 15.4m and 10.0m, respectively. Although the environments of relict forest trees showed a slight difference by species, the relative appearance frequencies of most trees were high in the environments where the altitude was higher than 1,000 m, slope degree was greater than $25^{\circ}$, the slope faced north, and microtopography was at the upper of slopes. Regarding the stand characteristics of relict forest trees per unit area ($/100m^2$), the average total coverage was 294% (max. 475%), the total average number of species was 36 species (max. 60 species), the average species diversity index (H') was 2.560 (max. 3.593), the average canopy closure was 84.8% (max. 94.6%), and the average basal area (/ha) was $52.7m^2$ (max. $116.4m^2$, relict trees $30.0m^2$, and other trees $22.7m^2$). The analysis of the dynamics of the forest stands where relict forest trees were growing showed four types of the maintenance mechanisms of relict forest trees depending on the supply pattern of succeeding trees: "Low-density but persistent type (Quercus mongolica, Abies holophylla, Tilia amurensis, and Pyrus ussuriensis)," "Long ago stopped type (Pinus densiflora)," "Recently stopped type (Abies nephrolepis, Quercus variabilis, and Betula schmidtii)," and "Periodically repeated types of supply and stop (Salix caprea and Quercus serrata).".

A Study of The planting Arrangement of Ornamental Trees And Shrubs in Intermane Buddhist Temples. (산지형 사찰에 있어 조경식물 배치형식에 관한 연구)

  • Shim, Jai-Sung;Bae, Jeong-Kwan;Seo, Byung-Key
    • The Journal of Natural Sciences
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    • v.14 no.1
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    • pp.63-81
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    • 2004
  • It is the purpose of this study to arrange in ornamental trees and shrubs the planting that leads to an appropriate type of templescape. The study was designed primarily as an examples for each Buddhist temple which desires further decoration with several ornamental plants, doubles the effectiveness of the Sen-Buddhist meditation, and boosts tourists' attraction. To investigate the planting status and type of trees and shrubs in the precincts of Buddhism temples, We close three temples : They are Nagsansa, Boolgoogsa and Booseugsa, which are intermonatane area temples all together. The results investigated were summarized as follows :1. Planting status and pattern in temples Open spaces of the Daewoogjeon in all temples, a main Sanctuaries in temple buildings, where Buddha is enshrined in, we could not find any kind of trees of shrubs to be planted.Muryangsujeon, a symbol of "Future", which can be also found at Boosugsa temple, is living in Western Elysium world and takes mercy on mankind of this life. Taxus cuspidata was planted at this Muryangsujeon, known as an immeasurable bliss building, where an Amitabha is enshrined in.Total 25 species of trees and shrubs were planted around Birozani building of Buddhist temples, Birozani is enshrined at the Birojion of Boollgoogsa temple.The buddhist Goddess of Mercy which is a buddhist saint for pursuit of fortune and blessing to relieve the mankind is enshrined at Wonchonjeon, Daebijeon and Kwaneumjeon which are able to observe at both Boolgoogsa and Nagsnsa temples, where Euonymus japonicus trees including other 26 species could be found in common at both temples.2. Correlation between trees/shrubs and temple buildingsTrees and flowers symbolizing Buddha are often planted as material sources of gardening to decorate : They are Logerstroemia indica as Buddha's flower, Viburmum opulus var. calvescens resembling Buddha's head, Tilia mandshurica producing the beads of rosary, Gardenia jaminoides Ellis with white flower blade and flower of bliss, not flower to this day and Lotus flowering clearly in the pond filling with dirty water which is able to clarifies the world full of crime, infidelity and injustification. Among these Buddhist' plants, however, Logerstroemia indica could be found in all three temples, and Viburmum opulus var. calvescens at both Nagsansa and Boosugsa. Also, Lager stroemia indica was planted at all three temples and Viburmum opulus var. calvescens at both temples of Nagsansa and Boosugsa. Tilia mandshurica and Gardenia jasminoides Ellis were not found in any temples which might become the subject of investigation.In relation of the buildings of each temples as a sanctified space, the planting of trees and shrubs was not considered for the arrangement, templescape architecture or species. And, also, we could not find in the study any special relationship of trees/ shrubs with the characteristics of temples.With the results obtained through precise studies we presented here in this paper newly designed model of templescape in intermane buddhist temple which can be applied for planting and arrangement of trees or/and shrubs. Basic principles of model in mind are:To consider the correlation of the dominant between plants and temple buildings.To plant trees/shrubs for special functions as well as conditions of temple location.To make tree arrangement correlating to Buddhism spirit.To induce environment friendly plants to be planted, suitable to regional conditions.This redecorated model of templescape might be used as a canon of the tree planting and arrangement in the precincts of Buddhism temples.

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A Study on the Application of IUCN Category to the Protected Areas of Korea (우리나라 보호지역에 IUCN 카테고리 적용 방안에 관한 연구)

  • Heo, Hag-Young;Kim, Hyun;Lee, Yeong-Joo;Kim, Seong-Il
    • Journal of Environmental Policy
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    • v.6 no.2
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    • pp.71-96
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    • 2007
  • This study aimed at effectively applying the IUCN category system to the protected areas in Korea. In addition, in order to change IUCN category of national parks to IUCN category II and to review the application of IUCN category classification key, a case study was conducted in Sobaeksan National Park. In order to apply the IUCN category system to the protected areas in Korea, a flexible approach appropriate to characteristics in Korea is required for management objectives of protected areas, including protection of wildemess, sustainable use of resources and preservation of cultural and traditional features. In addition, considerations of restrictions on use area and use districts, relative comparison of use types (visit, use of resources, residence) by IUCN category and use of combined classifications are necessary. Principles for the application of the IUCN category include (1)exclusion of wilderness protected areas (Ib), (2) extremely limited use regarding the sustainable use of natural resources(sum of natural preservation area and natural environment area is over 95%), (3) considerations of management conditions, including residential occupation level, (4) preservation of ecosystem services, and (5) use of combined classifications. In addition, in accordance with these principles, IUCN category classification key was suggested. When this was applied to the case study area, Sobaeksan National Park was classified as IUCN Category II and Taxus cuspidata community, which is designated as a natural monument, was classified to be Category Ia. Classification key suggested in this study may be used as basic data for applying categories in the future. Since detailed review on the practical improvement direction of laws and regulations and systematic alternatives, which are required before introducing IUCN category, are poor, in order to manage the protected areas efficiently by applying the IUCN category in the future, studies on management means appropriate to the conservation objectives of each category are necessary. This would allow management differentiated for each category.

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The Formative Characteristics of Seogo-jeongsa & Sameun-jeong Byeolseo Gardens in Toerori Miryang (밀양(密陽) 퇴노리(退老里) 서고정사(西皐精舍)와 삼은정(三隱亭) 별서(別墅)의 조영(造營) 특성(特性))

  • Lee, Hyun-Woo;Rho, Jae-Hyun
    • Journal of the Korean Institute of Traditional Landscape Architecture
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    • v.31 no.4
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    • pp.70-83
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    • 2013
  • To widen prospect of villa as Korea traditional garden, the result of the research about constructional characteristics of Seogo-jeongsa and Sameun-jeong of Yeoju Lee family at Toero-ri, Miryang-si, Gyeongsangnam-do, Korea, is abstracted as below. Seogo-jeongsa(西皐精舍) and Sameun-jeong(三隱亭) was intended to practise retirement with sense of unity with the head residence, since Seogo-jeongsa was 340m far and sameun-jeong was 630m far away from head residence, which set within 1km. Although Seogo-jeongsa's basic prop, Sameun-jeong's timber, which are both from designer's pen name and head residence name, "Hangjae(恒齋)" and "Yongjae(庸齋)," and Hanseoam have each different exposure, it is homogeneous as space configurational side in order to optimize the gaze leading effect and appreciated domestic trees from the pond, which is center of the garden and the significant facility. Method of direction of Seogo-jeongsa's Hwalsudang(活水塘) and Sameunjeong's pond of Sameun-ji(三隱 池) gets attention by distinction. Especially, directing of 3 step water flow from behind to front and traditional pond method, called Bangji-Bangdo(方池方島), were very unique that any other place can be found. Also, the middle islet on the pond at both villa, which isn't more supper, but more for the symbolic molding that seek a stone pagoda(石假山), is more interesting directing method as considering the relationship of the 3 stone pagoda imageries on the surface of Ghwayukchon(果肉泉) At the same time, Seogo-jeongsa and Sameun-jeong shows very different characteristics from traditional villa that adopted both domestic trees and foreign trees. Particularly, it is extremely exceptional case to plant vigorous needleleaf tree, such as Chamaecyparis pisifera, C. obtusa, Pinus bungeana, P. palustris, Sciadopitys verticillata, Metasequoia glyptostroboides and Cedrus deodara, on Sameun-jeong. Moreover, adopting foreign wild needleleaf trees for landscaping trees, such as a Torreya nucifera, Taxus cuspidata, P. parviflora, and foreign landsacping trees, such as P. bungeana, Cryptomeria japonica and C. obtusa tells planting trend of the late Joseon dysnasty era. Also, as we can know from 2 Jipgyeong(集景), which is 'Seogo-jabyoung 17 young(西皐雜詠十七詠)', and 'Sameun-jeong 12 Gyoung(三隱亭十二景)' which are set on both villa, the intend to expand the garden area is strongly shown by the natural forest directing. As a result, Seogo-jeongsa and Sameun-jeong, located at Toero-li Miryang, are sharing the traditional Joseon dynasty era's custom, such as space and visual composition; however, it is different and attractive garden remains as a point of view of water directing, stone pagodas, and adopting foreign landscaping trees.

Floristic features of upland fields in South Korea (우리나라 밭 경작지에 출현하는 식물상 특성)

  • Kim, Myung-Hyun;Eo, Jinu;Kim, Min-Kyeong;Oh, Young-Ju
    • Korean Journal of Environmental Biology
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    • v.38 no.4
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    • pp.528-553
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    • 2020
  • Upland fields are characterized by dry environments, a high degree of disturbance by farming practices such as double-cropping, and a high diversity of crops compared to other field types. This study focused on the floristic composition and characteristics of upland fields in South Korea. Flora surveys were conducted in 36 areas in nine provinces at two times (June and August) in 2015. The results showed that the vascular plants in the upland fields in South Korea included 532 taxa, containing 100 families, 322 genera, 483 species, nine subspecies, 37 varieties, one form, and two hybrids. Among the 100 families, Asteraceae was the most diverse in species (75 taxa), followed by Poaceae (68 taxa), Fabaceae (34 taxa), Polygonaceae (21 taxa), Rosaceae (19 taxa), and Liliaceae (17 taxa). Based on the occurrence frequency of each species, Acalypha australis L. (100%), and Artemisia indica Willd. (100%) were the highest, followed by Humulus scandens (Lour.) Merr., Rorippa palustris (L.) Besser, Conyza canadensis (L.) Cronquist, Erigeron annuus (L.) Pers., Lactuca indica L., Commelina communis L., Digitaria ciliaris (Retz.) Koeler, Echinochloa crus-galli(L.) P.Beauv., Cyperus microiria Steud., and Oxalis corniculata L. The biological type of upland fields in South Korea was determined to be Th-R5-D4-e type. Rare plants were found in 11 taxa: Taxus cuspidata Siebold & Zucc, Magnolia kobus DC, Clematis trichotoma Nakai, Aristolochina contorta Bunge, Buxus sinica (Rehder & E.H.Wilson) M.Cheng var. koreana (Nakai ex Rehder) Q.L.Wang, Melothria japonica (Thunb.) Maxim, Mitrasacme indica Wight, Lithospermum arvense L., Carpesium rosulatum Miq., Allium senescens L., and Pseudoraphis sordida (Thwaites) S.M.Phillips & S.L.Chen. Ninety-seven taxa contained naturalized plants composed of 24 families, 68 genera, 97 species, one variety, and one form. The urbanization and naturalization indices were 30.5% and 18.4%, respectively.

A study on the ecological habitat and protection of natural Sorbus commixta forest at Mt. Seorak (설악산(雪嶽山)에 분포(分布)하는 마가목 천연림(天然林)의 생태환경(生態環境)과 보호(保護)에 관(關)한 연구(硏究))

  • Shin, Jai Man;Kim, Tong Su;Han, Sang Sup
    • Journal of Forest and Environmental Science
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    • v.3 no.1
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    • pp.1-9
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    • 1983
  • The purpose of this study was to elucidate the ecophysiological habitat of natural Sorbus commixta forest at Mt. Seorak. The results obtained were as follows: 1. The Sorbus commixta trees mainly distributed from 900m to 1,500m altitude. In there, the warm index(WI) was about 42$3.2{\times}10^3$ to $9.2{\times}10^3$, cation exchange capacity(CEC) was 13.7 to 19.5mg/100g, N content 0.21 to 0.39%, $P_2O_5$ content was 22.6 to 38.7ppm, and pH value was 5.6 to 5.8 respectively. 4. The upper crown trees in Sorbus commixta communities were Abies nephrolepis, Taxus cuspidata, Betula platyphylla var. japonica, Quercus${\times}$grosseserrata, Acer mono, Prunus sargentii, Carpinus cordata, Tilia amurensis, and the under crown trees were Rhododendron brachycarpum, Acer pseudo-sieboldianum, Thuja olientalis, Corylus heterohpylla, Philadelphus schrenckii, Rhododendron schlippenbachii, Rhododendron mucronulatum, and Magnolia sieboldii. 5. The stand densities were 1,156 trees/ha at 1,160m and 3,600 trees/ha at 1,300m respectively. The coverages by the DBH basal area were 0.37 at 1,160m and 0.31 at 1,300m respectively, and the vegetation coverages by the crown projection area were 2.04 at 1,160m and 1.61 at 1,300m respectively. 6. The light extinction coefficient(k) in Beer-Lambert's law, showed the distance, F(z), from top canopy to aboveground, was 0.17. 7. The water relations parameters of Sorbus commixta shoot were obtained by the pressure chamber technique. The osmotic pressure, ${\pi}_o$, at maximum turgor was -16.2 bar, and VAT pressure was 14.5bar. The osmotic pressure, ${\pi}_p$, at incipient plasmolysis was -19.4bar. The relative water contents at incipient plasmolysis were 83.1% ($v_p/v_o$) and 87.1%($v_p/w_s$;$w_s$, total water at maximum turgor). 8. The bulk modulus of elasticity(E) of shoot was about 69.6. The total symplasmic water to total water in shoot was 67.7%, and the apoplastic water to total water was 32.3%.

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