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Agronomic Characteristics of Common Millet Germplasm (기장 유전자원의 농업적 특성)

  • Yoon, Seong-Tak;Xu, Zhen Yu;Kim, Seong-Min;Kim, Chang-Yung
    • KOREAN JOURNAL OF CROP SCIENCE
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    • v.53 no.4
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    • pp.394-400
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    • 2008
  • Crop characteristics of 137 common millet (Panicum miliaceum L.) resources collected in Korea were investigated in order to establish basic data for the improvement of crop breeding. Spike types of 137 plant resources were classified as 4 types of oblate, spreading, oblate-compact (intermediary type between oblate and compact) and oblate-spreading (intermediary type between oblate and compact) type, of which oblate spike type was the highest percentage of 66.4% (91 plant resources) of 137 germplasm. Seed color was classified as 5 colors of dark brown, yellow, brown, yellowish brown, and yellowish white, of which yellowish white was the highest percentage of 66.4% (54 plant resources) among them. Seed type was classified as 2 types of pyramid shape and short pyramid shape, which showed 62.8% and 37.2% respectively. Days from seeding to heading date showed the range from 68 to 85 days with 73.7 days in the average and the highest frequency proportion of it was the group from 70 to 75 days, which occupied 46.7% (64 plant resources) of 137 germplasm. Days from seeding to physiological maturity showed the range from 109 to 124 days with 112.0 days in the average and the highest frequency proportion (60.6%) of it was the group from 111 to 115 days, while there were also occupied with 6.6% below 110 days and 5.1% over 120 days among 137 germplasm. Group from 181 cm to 190 cm of culm length showed the highest frequency proportion of 47.4%, whereas below 160 cm was the lowest occupancy of 2.9%. Spike length showed the range from 181 to 190 cm with 181.4 cm in the average and the highest frequency distribution of it was the group from 31 to 35 cm with the proportion of 54.0%. Number of grains per spike showed the range from 432 to 2,665 grains with 1,429.5 grains in the average and group from 1,200 to 1,400 grains per spike was the highest frequency distribution with the proportion of 20.4%. There were also occupied with 13.1% below 1,000 grains and 8.6% over 2,000 grains per spike among 137 germplasm.

Studies on Growth, Forage Yield, and Nutritive Value according to Different Seeding Dates of Barnyard Millet (파종기에 따른 사료용 피의 생육, 수량 및 사료가치에 관한 연구)

  • Lee, Jung-Joon;Kim, Jung-Gon;Sung, Byung-Ryul;Song, Tae-Hwa;Park, Tae-Sun
    • Journal of The Korean Society of Grassland and Forage Science
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    • v.33 no.4
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    • pp.245-251
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    • 2013
  • These experiments are being conducted to obtain data of plant height, panicle length, heading date, number of culm, and fresh and dry weight, crude protein, acid detergent fiber (ADF), neutral detergent fiber (NDF), total digestible nutrients (TDN) by six times seeding of 10 days interval ranging from May 1st to June 21st for two types of barnyard millet from the Suwon area in Korea. In the early-heading type, IT 170609 (Echinochloa crus-galli), the plant height and the number of culm per square meters are decreased by late seeding relatively, and heading date is 7 to 10 days earlier than the late seeding. And the dry weight of forage is rapidly decreased after the June 11th seeding. In the late-heading type, IT 195422 (Echinochloa frumentacea), the plant height and the number of culm per square meters are also decreased by late seeding relatively, and heading date is about 13 days earlier than late seeding. The dry weight of forage was rapidly decreased after the June 11th seeding. In both two types, the nutritive value of ADF, NDF, and TDN are not changed by late seeding, but crude protein is increased by late seeding. The TDN yields of late-heading type, IT 195400, show average amounts in the June 1st seeding, but the TDN yield of early-heading type, IT 170606, indicate safety amounts of dry weight to the March 21st seeding. And these data are available for cropping systems, cultivating barnyard millet in summer and cereals for forage in winter, to get high production of forage in Korea.

Characteristics of Seepage Water and Groundwater in a Coastal LPG Storage Cavern of Jeonnam (전남 해안 LPG 저장공동 유출수와 주변 지하수의 수질특성)

  • Lee, Jin-Yong;Choi, Mi-Jung;Kim, Hyun-Jung;Cho, Byung-Wook
    • Journal of Soil and Groundwater Environment
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    • v.14 no.4
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    • pp.33-44
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    • 2009
  • Water curtain of an underground LPG storage cavern is a facility to prevent leakage of high pressure gases, for which groundwater should flow freely towards the cavern and groundwater level also must be stably maintained. In this study, in order to evaluate qualities of seepage water and surrounding groundwater of an underground LPG storage cavern in Yeosu, 4 rounds of samplings, field measurements and laboratory analyses (February, May, August, October of 2007) were conducted. According to field measurements, pH was weak acidic to neutral but it gradually increased with time. Electrical conductivity (EC) of groundwater near a salt stack showed very high values between 10.47 and 38.50 mS/cm. Dissolved oxygen (DO) showed a very wide range of 0.20~8.74 mg/L and a mean of oxidation-reduction potential (ORP) was 159 mV, which indicated an oxidized condition. Levels of $Fe^{2+}$ and $Mn^{2+}$ were mostly less than 3 mg/L. All of seepage waters showed a Na-Cl type while only groundwater near the salt stack showed a Na-Cl type with a high total dissolved solid. The other groundwaters exhibited typical $Ca-HCO_3$ types. Levels of aerobic bacteria were mostly very high (573-39,520 CFU/mL). Based on the analyses of these hydrochemistry and biological characteristics, it is concluded that there are no particular problems in groundwater and seepage water, which not causing a trouble in the cavern operation. However, both for control of bio-clogging and for sustainable operation of the water curtain system, a regular hydrochemical and microbiological monitoring is required for the seepage water and surrounding groundwater.

Effects of Forage-Rice Cropping Systems on the Growth and Grain Quality of Early Maturing Rice Cultivars and Soil Chemical Properties in Paddy Fields in Southern Korea (사료작물-벼 작부체계가 조생종 벼의 생육과 미질 특성 및 토양의 화학적 특성에 미치는 영향)

  • Oh, Seo Young;Oh, Seong Hwan;Seo, Jong Ho;Choi, Jisu
    • KOREAN JOURNAL OF CROP SCIENCE
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    • v.66 no.4
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    • pp.297-306
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    • 2021
  • To select rice (Oryza sativa L.) cultivars suitable for forage-rice double cropping system, the growth and grain quality of four early maturing rice cultivars (Joun, Jopyeong, Haedamssal, and Unkwang), and the chemical properties of soils were investigated under single- (fallow-rice) and forage-rice double-cropping systems in paddy fields in Miryang, southern Korea. The soil where two forage crops [Italian ryegrass (Lolium multiflorum Lam.) and oat (Avena sativa L.)] were cultivated during winter had a slightly lower pH; an increase in total nitrogen (T-N), K, Ca, and Na contents; and a slight decrease in organic matter and available P2O5 contents, compared with the soil fallowed during winter. This shows that the chemical properties of paddy soils can be improved by winter forage cropping. At the heading stage, the culm length, panicle length, panicle number, and leaf color of all cultivars, except for Haedamssal, were generally higher under double-cropping than under single-cropping. For Haedamssal, the culm length and leaf color did not differ between the cropping systems, but the panicle length was slightly shortened and its panicle number increased under double-cropping. After harvest, the yield of milled rice decreased for all cultivars except Haedamssal, but increased in Haedamssal under double-cropping. The head rice rate was slightly higher under double cropping, particularly in Jopyeong and Haedamssal, than under single-cropping. The protein content of milled rice under double cropping was higher and its amylose content was similar or slightly lower compared to those of rice under single cropping, resulting in decreased Toyo values for rice under double-cropping. The pasting temperature did not differ significantly between the cropping systems. However, Haedamssal had a low pasting temperature but a high Toyo value under double cropping, compared to the other three cultivars, suggesting that its palatability is relatively high. Furthermore, panicle number increased and milled rice yield did not decrease, even under double cropping. Therefore, Haedamssal seems to be the best cultivar for paddy-based double cropping with forage crops.

Improving the Current Status and Cultural Value of Donguibogam Wanyoung Woodblocks (<동의보감(東醫寶鑑)> 완영(完營)책판의 현황과 문화재적 가치 제고(提高))

  • KIM, Hwaseon
    • Korean Journal of Heritage: History & Science
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    • v.55 no.2
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    • pp.50-64
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    • 2022
  • This thesis is a study on the woodblock of Donguibogam by Jeolla Gamyeong(Wanyoung Woodblock). It was registered as a tangible cultural property in Jeollabuk-do in 2005. The purpose of this article is to determine the current status and value of the Donguibogam Wanyoung Woodblock. Jeonju Hyanggyo's Wanyoung Woodblock was used to print books in Gamyeong, Jeolla Province. Currently, ten kinds of woodblocks are preserved. In 1987, a wooden bookshelf was installed and preserved in "Jangpangak" of Jeonju Hyanggyo. It is now preserved and managed at Chonbuk National University Museum. Among the ten woodblocks, the Wanyoung woodblock of Donguibogam is presumed to be a woodblock of a book printed by Jeolla Gamyeong in 1814. The value of Donguibogam has long been recognized domestically and internationally. Donguibogam, compiled in 1610, was first published in 1613 in the wood type of Gaeju Gapinja. Following its publication, its value was recognized not only in Joseon but also in China and Japan. In 2009, the first edition of Donguibogam was registered as a UNESCO World Heritage. Accordingly, Donguibogam has been recognized for various aspects, but little research has been conducted on the woodblocks that printed Donguibogam. Therefore, this paper analyzes the current status of the Wanyoung woodblock of Donguibogam, aiming to enhance its meaning and value. King Seonjo presented the ideals of public health care and preventive medicine when compiling Donguibogam, which was distributed according to his instructions. For this reason, the first edition of Donguibogam was registered as a UNESCO World Heritage. It can be said that the production of Donguibogam woodblocks was an important tool in realizing Joseon's ideals through national dissemination. Furthermore, the woodblock of Donguibogam represents the spirit of thinking about the people, going beyond the purpose of printing, and it was meaningful in the spread of medical knowledge among the people. In this article, I will examine the overall contents of the Wanyoung woodblock of Donguibogam to enhance its meaning and value. The results show that the Wanyoung woodblock of Donguibogam is meaningful in that it is almost the only one produced and stored by Gamyeong. Moreover, Wanyoung woodblocks are meaningful in that perfect editions could be preserved and disseminated for a long time, unlike other wood-type prints.

Farmer's Field Trial of Different Coating and Covering Materials on Rice Growth and Yield in Wet Hill Seeded Rice (볍씨 코팅 및 규산복토에 따른 벼 무논점파재배 농가실증시험연구)

  • Park, Kwang Ho;Kim, Yang Sik;Chang, Jin Tack
    • Journal of Practical Agriculture & Fisheries Research
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    • v.17 no.1
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    • pp.85-92
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    • 2015
  • The experiment was conducted to evaluate rice growth and yield as affected by different coating and covering materials such as a iron, silicate, iron and silicate mixture of rice seeds in farmer's rice growing field. The tiller number was 36.7 at iron-coated seeds, 32.8 at silicate-covered seeds, 30.3 at iron and silicate mixture coated seeds and 30.2 at untreated control in 44days after seeding. The seedling height was 38.2cm of iron and silicate mixture, 37.7cm of untreated control, 36cm of iron-coated and 35.7cm of silicate covered seeds in 43days after seeding. At 75days after seeding rice tiller number was 153 of iron-coated seeds, 152 of silicate-covered seeds, 147 of untreated seeds and 141 of iron and silicate mixture-coated seeds and also there were different plant height growth of 87.4cm in silicate-covered seeds, 85.7cm in iron and silicate mixture, 85.4cm in untreated control and 83.0cm in iron-coated seeds. The panicle length was of 21.0cm in iron and silicate mixture coated seeds, 20.8cm in silicate covered seeds, 20.7cm in untreated control seeds and 20.6cm in iron-coated seeds. The panicle number was 464 at iron-coated seeds, 404 at untreated control seeds, 427 at silicate-covered seeds and 412 at iron and silicate mixture coated seeds. The spikelet number per m2 was of 32,503 in iron-coated seeds, 31,813 in silicate-covered seeds, 29,646 in untreated control, 28,896 in iron and silicate mixture coated seeds. The ripened ratio of rice grain was of 94.5% at iron-coated seeds, 93.9% at iron and silicate mixture coated seeds, 93.6% at silicate covered seeds and 93.2% at untreated control seeds. The rice yield was of 591kg/10a at iron-coated seeds, 580kg/10a at silicate-covered seeds, 571kg/10a at iron and silicate mixture-coated seeds and 539kg/10a at untreated control.

Growth and Yield in Direct Seeded Rice Cultivation with Iron Coated-Seeds (철분코팅 볍씨를 이용한 벼 직파재배의 생육 특성 및 수량)

  • Park, K.H.;Park, S.T.
    • Journal of Practical Agriculture & Fisheries Research
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    • v.20 no.1
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    • pp.5-18
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    • 2018
  • The field trial was performed to evaluate the rice growth and yield in direct seeding cultivation with iron-coated rice seeds. The required time for seed emergence was for 9~11days in the tested direct seeding methods. That was 1~2days earlier in direct seeding with pregerminated seeds than that of direct seeding with iron-coated seeds. The seedling establishment was highest in water seeding with iron-coated seeds but there was not significant difference in terms of statistical analysis. The rice plant height was taller in water seeding with broadcasting method than that of wet hill-seeding methods and in direct seeding with iron-coated seeds than that of direct seeding with pregerminated seeds. The tiller number in the rice plant was the highest in machine transplanting at 30days after direct seeding(June 17) and in water seeding with iron-coated seeds at 45days after seeding(DAS) and 60DAS. The tiller number of 75 and 90DAS in the tested rice cultivation methods being with 352~405/m2 was not significantly different in terms of statistical analysis. The heading time was not different in rice direct seeding methods but 2 day earlier in direct seeding with iron-coated seeds than that of direct seeding with pregerminated seeds. The culm length was the highest in water seeding with iron-coated seeds and the panicle length was the longest in wet hill-seeding with pregerminated seeds. The panicle number per m2 was highest in water seeding with iron-coated seeds but not significant difference among the tested rice cultivation methods. The water seeding with iron-coated seeds resulted in the highest spikelet number per m2 and the heaviest grain weight of brown rice. Percentage of ripened kernel was the highest in wet hill-seeding with iron-coated seeds. But there were not significant among the tested rice cultivation methods. The milled rice yield in direct seeding methods was 3~21% higher than that in machine transplanting. Water seeding with iron-coated seeds recorded the highest milled rice yield being with 6.86t/ha.The occurrence of sheath blight was high according to machine transplanting>wet hill-seeding>water seeding. Weed occurrence was the highest in water seeding with pregerminated seeds. Weedy rice occurred not in machine transplanting but occured 0.6~0.7% in direct seeding methods with pregerminated seeds and 0.1% in direct seeding with iron-coated seeds.

Evaluation of Genetic Safety in Genome-editing Rice Through Comparative Analysis of Genetic and Agronomic Traits (유전적 특성과 농업형질의 비교분석을 통한 유전자 교정 벼의 안전성 평가)

  • Seung-Kyo Jeong;Dohyeong Gwon;Bae-Hyeon Lee;Jeong-Hwan Suh;Rahmatullah Jan;Jae-Ryoung Park;Taehun Ryu;Kyung-Min Kim
    • Journal of Life Science
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    • v.34 no.8
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    • pp.567-575
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    • 2024
  • New breeding techniques (NBT) recognize specific DNA sequences and remove, modify, or insert DNA at a desired location, and can be used to treat genetic diseases in humans or to improve the traits of livestock or crops. In this study, we conducted a comparative analysis of various agricultural traits and assessed the safety of gene transferability in third-generation genome-editing rice (OsCKq1-G3) with T and G nucleotide insertions developed using the CRISPR/Cas9 SDN-1 method, in comparison to its parental line (Oryza sativa L., cv Ilmi). The analyzed traits included heading date, culm length, panicle length, tiller number, yield, germination rate, viviparous germination rate, shattering, after wintering seed viability, the presence of toxins and allergens. The target trait, heading date, exhibited a high significant difference of approximately 5 days. Culm length, panicle length, tiller number, yield showed no significant differences compared to the parental line. No T-DNA bands indicating gene transfer were detected. In the third generation of genome-edited rice, the T-DNA was confirmed to be eliminated as successive generations advanced through self-pollination. Through the analysis of germination rate, viviparous germination rate, shattering, and after wintering viability, we confirmed that the genome-editing rice has no potential for weediness. The ORF and amino acid sequences of the genome-editing rice did not reveal any toxins and allergens. The results of this study can be utilized as important data for the environmental risk assessment of genome-editing rice.

Genetic Analysis of Quantitative Characters of Rice (Oryza sativa L.) by Diallel Cross (이면교배(二面交配)에 의한 수도량적(水稻量的) 형질(形質)의 유전분석(遺傳分析)에 관(關)한 연구(硏究))

  • Jo, Jae-seong
    • Korean Journal of Agricultural Science
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    • v.4 no.2
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    • pp.254-282
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    • 1977
  • To obtain information on the inheritance of the quantitative characters related with the vegetative and reproductive growth of rice, the $F_1$ seeds were obtained in 1974 from the all possible combinations of the diallel crosses among five leading rice varieties : Nongbaek, Tongil, Palgueng, Mangyeong and Gimmaze. The $F_1$'s including reciprocals and parents were grown under the standard cultivation method at Chungnam Provincial Office of Rural Development in 1975. The arrangement of experimental plots was randomized block design with 3 replications and 12 characters were used for the analysis. Analytical procedure for genetic components was followed the Griffing's and Hayman's methods and the results obtained are summarized as follows. 1. In all $F_1$'s of Tongil crosses, the longer duration to heading was due to dominant effect of Tongil and each $F_1$ showed high heterosis in delaying the heading time. It was assumed that non-allelic gene action besides dominant gene effect might be involed in days to heading character. However, in all $F_1$'s from the crosses among parents excluding Tongil the shorter duration was due to dominant gene action and the degree of dominance was partial, since dominance effects were not greater than the additive effect. The non-allelic gene interaction was not significant. Considering the results mentioned above, it was regarded that there were two kinds of Significantly different genetic systems in the days to heading. 2. The rate of heterosis was significantly different depending upon the parents used in the crosses. For instance, the $F_1$'s from Togil cross showed high rate of heterosis in longer culm. Compared to short culm, longer culm was due to recesive gene action and short culm was due to recesive gene action. The dominant gene effect was greater than the additive gene effect in culm length. The narrow sense of heretability was very low and the maternal effects as well as reciprocal effects were significantly recognized. 3. The lenght of the of the uppermost internode of each $F_1$ plant was a little lorger than these of respective parental means or same as those of parents having long internodes, indicating partial dominance in the direction of lengthening the uppermost internodes. The additive gene effects on the uppermost internode was greater than the dominance gene effect. The narrow as well as broad sense of heritabilities for the character of the uppermost internode were very high. There were significant maternal and reciprocal effect in the uppermost internode. 4. The gene action for the flag leaf angle was rather dominance in a way of getting narrower angle. However, in the Palgueng combinations, heterosis of $F_1$ was observed in both narrow and wide angles of the flag leaf. The dominant effects were greater than the additive effects on the flag leaf angle. There were observed also a great deal of non-allelic gene interacticn on the inheritance of the flag leaf angle. 5. Even though the dominant gene action on the length and width of flag leaf was effective in increasing the length or width of the flag leaf, there were found various degrees of hetercsis depending upon the cross combination. Over-dominant gene effect were observed in the inheritance of length of the flag leaf, while additive gene effects was found in the inheritance of the width of the flag leaf. High degree of heretabilities, either narrow or broad sense, were found in both length and width of the flag leaf. No maternal and reciprocal effect were found in both characters. 6. When Tongil was used as one parent in the cross, the length of panicle of $F_1$'s was remarkedly longer than that of parents. In other cross comination, the length of panicle of $F_1$'s was close to the parental mean values. Rather greater dominent gene effect than additive gene effect was observed in the inheritance of panicle length and the dominant gene was effective in increasing the panicle length. 7. The effect of dominant genes was effective in increasing the number of panicles. The degree of heterosis was largely dependent on the cross combination. The effect of dominant gene in the inheritance of panicle number was a little greater than that of additive genes, and the inheritance of panicle number was assumed to be due to complete dominant gene effects. Significantly high maternal and reciprocal effects were found in the character studied. 8. There were minus and plus values of heterosis in the kernel number per panicle depending upon the cross combination. The mean dominant effect was effective in increasing the kernel number per panicle, the degree of dominant effect varied with cross combination. The dominant gene effect and non-allelic gene interaction were found in the inheritance of the kernel number per panicle. 9. Genetic studies were impossible for the maturing ratio, because of environmental effects such as hazards delaying heads. The dominant gene effect was responsible for improving the maturing ratio in all the cross combinations excluding Tongil 10. The heavier 1000 grain weight was due to dominant gene effects. The additive gene effects were greater than the dominant gene effect in the 1000 grain weight, indicating that partial dominance was responsible for increasing the 1000 grain weight. The heritabilites, either narrow or broad sense of, were high for the grain weight and maternal or reciprocal effects were not recognized. 11. When Tongil was used as parent, the straw weight was showing high heterosis in the direction of increasing the weight. But in other crosses, the straw weight of $F_1$'s was lower than those of parental mean values. The direction of dominant gene effect was plus or minus depending upon the cross combinations. The degree of dominance was also depending on the cross combination, and apparently high nonallelic gene interaction was observed.

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The Effect of Mixed Cultivation of Rice Varieties on the Plant Characteristics and the Occurrence of Blast under Various Fertilizer Levels (혼식(混植)한 몇가지 수도품종(水稻品種)의 생육(生育)과 병해발생(病害發生))

  • Lee, Kwang Soo;Ahn, Su Bong
    • Korean Journal of Agricultural Science
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    • v.14 no.2
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    • pp.232-247
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    • 1987
  • Three rice varietis, Dongjinbyeo, Daecheongbyeo and Chucheongbyeo, which are leading cultivars in the Chungnam area, were planted to determine their effects of mixture on the plant characteristics and the blast occurrence. The summary obtained is as follows; 1. The heading dates of each cultivars were not affected by the mixing culture. Only a day was delayed in heading date of Chucheongbyeo under commonly standard fertilizer level. 2. The effects of cultivar mixture on the plant height was high when low level of fertilizer were applied. Of the cultivar mixture, the mixing of Dongjinbyeo and Daecheongbyeo shoed 5.1% of plant height increase. The effects of mixture on the tillering number were apparent only under the standard fertilizer level. The mixture of Dongjinbyeo and Chucheongbyeo was effective in increasing the tillering number up to 5.7% over mean tillering number of each cultivars. 3. The stem height tended to increase as the fertilizer level increased and the effects of cultivar mixture on the stem height was high. The stem height of mixture of Dongjinbyeo and Daecheongbyeo was highest compared with other mixture cropping. On the contrary, the panicle length was shortened as the fertilizer level decreased and cultivars were mixed. Among the cultivar mixtures, the mixture of Dongjinbyeo and Daecheongbyeo showed the most effectives of increasing the panicle length. 4. The total dry weight of mixed and pure cultivars were compared. The mixture of two cultivars was effective in increasing the dry weight of rice upto 3.5% over the average of dry weight of each cultivars. When three cultivars were mixed, there were 8.5% of dry weight increase over not mixed cultivars. The effects of mixture on the dry weight were more apparent under standard fertilizer level. 5. The rates of Neck and Node and Branch Blast occurrence were reduced when cultivars were mixed, and their reduction rates were 11.7% and 14.0%, respectively. The occurrence of Blasts was also decreased under the lowest fertilizer level than the standard fertilizer level. The least Blast occurrence was obtained when Dongjinbyeo and Daecheongbyeo were mixed cropped. 6. Significantly high number of spikelets of per square meter was observed when Chucheongbyeo and Dongjinbyeo were mixed. However, the spikelets of panicle were the lowest when Chucheongbyeo and Dongjinbyeo were mixed. The number of panicles per square meter was increased as the fertilizer levels increased, while the number of spikelets per panicle increased as the fertilizer level decreased. 7. The effects of mixture on the maturing were apparent regardless of the fertilizer level. The 1000 grain weight was higher when lower level of fertilizer was applied. The effects of cultivar mixture on the 1000 grain weight were more apparent in the plots of Chucheongbyeo and Dongjinbyeo mixture and three cultivar mixture. 8. The grain yield increased when cultivars mixed. The increment under mixing cropping was 4.6% over mono cropping. The effects of cultivar mixture on the yield increase were more apparent under lower levels of fertilizer application. The highest yield increase was obtained when Chucheongbyeo and Daecheongbyeo were mixed, and the rate of yield increase was 6.8% over mono-cropping. 9. The grain yield was highly correlated with number of panicles per square meter and dry weight. There was not any significant relationships found among grain yield, spikelets of panicle and ripening percentage.

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