Acknowledgement
We are grateful to the Director and Dean of the Wildlife Institute of India for their help. We are grateful to the Rajasthan Forest Department, as well as the reserve officials and field employees at the RTR for granting permission and allowing our work, as well as for their assistance. Special thanks go to the RTR's nature guides and field helpers, M. S. Gurjar and S. Sharma, for their assistance. The Ministry of Environment and Forests, Government of India provided financial support through the Training Research and Academic Council.
References
- Andersen, R., Duncan, P., and Linnell, J.D.C. (1998). The European Roe Deer: The Biology of Success. Scandinavian University Press.
- Andheria, A.P., Karanth, K.U., and Kumar, N.S. (2007). Diet and prey profiles of three sympatric large carnivores in Bandipur Tiger Reserve, India. Journal of Zoology, 273, 169-175. https://doi.org/10.1111/j.1469-7998.2007.00310.x
- Athreya, V. (2006). Is relocation a viable management option for unwanted animals? - The case of the leopard in India. Conservation and Society, 4, 419-423.
- Avinandan, D., Sankar, K., and Qureshi, Q. (2008). Prey selection by tigers (Panthera tigris tigris) in Sariska Tiger Reserve, Rajasthan, India. Journal of the Bombay Natural History Society, 105, 247-254.
- Bagchi, S., Goyal, S.P., and Sankar, K. (2003). Prey abundance and prey selection by tigers (Panthera tigris) in a semi-arid, dry deciduous forest in western India. Journal of Zoology, 260, 285-290. https://doi.org/10.1017/S0952836903003765
- Balme, G.A., Hunter, L., and Braczkowski, A.R. (2012). Applicability of age-based hunting regulations for African leopards. PloS One, 7, e35209. https://doi.org/10.1371/journal.pone.0035209
- Borah, J., Sharma, T., Das, D., Rabha, N., Kakati, N., Basumatary, A., et al. (2014). Abundance and density estimates for common leopard Panthera pardus and clouded leopard Neofelis nebulosa in Manas National Park, Assam, India. Oryx, 48, 149-155. https://doi.org/10.1017/S0030605312000373
- Carbone, C., and Gittleman, J.L. (2002). A common rule for the scaling of carnivore density. Science, 295, 2273-2276. https://doi.org/10.1126/science.1067994
- Champion, F.W. (1928). With a Camera in Tiger-land. Doubleday, Doran & Company.
- Champion, H.G., and Seth, S.K. (1968). A Revised Survey of the Forest Types of India. Manager of Publications.
- Chapron, G., Miquelle, D.G., Lambert, A., Goodrich, J.M., Legendre, S., and Clobert, J. (2008). The impact on tigers of poaching versus prey depletion. Journal of Applied Ecology, 45, 1667-1674. https://doi.org/10.1111/j.1365-2664.2008.01538.x
- Chauvenet, A.L., Durant, S.M., Hilborn, R., and Pettorelli, N. (2011). Unintended consequences of conservation actions: managing disease in complex ecosystems. PloS One, 6, e28671. https://doi.org/10.1371/journal.pone.0028671
- Chundawat, R.S., Gogate, N., and Johnsingh, A.J.T. (1999). Tigers in Panna: preliminary results from an Indian tropical dry forest. In J. Seidensticker, S. Christie, and P. Jackson (Eds.), Riding the Tiger: Tiger Conservation in Human-dominated Landscapes (pp. 123-129). Cambridge University Press.
- Datta, A., Anand, M.O., and Naniwadekar, R. (2008). Empty forests: large carnivore and prey abundance in Namdapha National Park, north-east India. Biological Conservation, 141, 1429-1435. https://doi.org/10.1016/j.biocon.2008.02.022
- Edgaonkar, A., Chellam, R., and Qureshi, Q. (2007). Ecology of the Leopard (Panthera pardus fusca) in Satpura National Park and Bori Wildlife Sanctuary. Wildlife Institute of India.
- Efford, M. (2004). Density estimation in live-trapping studies. Oikos, 106, 598-610. https://doi.org/10.1111/j.0030-1299.2004.13043.x
- Efford, M.G. (2010). Secr: spatially explicit capture recapture in R. Department of Zoology, University of Otago.
- Efford, M.G. (2011). Estimation of population density by spatially explicit capture-recapture analysis of data from area searches. Ecology, 92, 2202-2207. https://doi.org/10.1890/11-0332.1
- Efford, M.G., Borchers, D.L., and Byrom, A.E. (2009). Density estimation by spatially explicit capture-recapture: likelihoodbased methods. In D.L. Thomson, E.G., Cooch and M.J., Conroy (Eds.), Modeling Demographic Processes In Marked Populations (pp. 255-269). Springer.
- Glanz, W.E. (1982). The terrestrial mammal fauna of Barro Colorado Island: censures and long-term changes. In E.G. Jr Leigh, A.S. Rand, and D.M. Windson (Eds.), The ecology of a tropical forest: seasonal rhythms and long-term changes. (pp. 455-468). Smithsonian Institution Press.
- Gray, T.N.E., and Prum, S. (2012). Leopard density in postconflict landscape, Cambodia: evidence from spatially explicit capture-recapture. The Journal of Wildlife Management, 76, 163-169. https://doi.org/10.1002/jwmg.230
- Harihar, A., Pandav, B., and Goyal, S.P. (2011). Responses of leopard Panthera pardus to the recovery of a tiger Panthera tigris population. Journal of Applied Ecology, 48, 806-814. https://doi.org/10.1111/j.1365-2664.2011.01981.x
- Henschel, P., Hunter, L., Breitenmoser, U., Purchase, N., Packer, C., Khorozyan, I., et al. (2008). Panthera pardus. In The IUCN Red List of Threatened Species v. 2015.2. Retrieved Apr 13, 2023 from https://www.iucnredlist.org.
- Hijmans, R.J., Cameron, S.E., Parra, J.L., Jones, P.G., and Jarvis, A. (2005). Very high resolution interpolated climate surfaces for global land areas. International Journal of Climatology, 25, 1965-1978. https://doi.org/10.1002/joc.1276
- Jhala, Y.V., Gopal, R., and Qureshi, Q. (2008). Status of the Tigers, Co-predators, and Prey in India. National Tiger Conservation Authority, Govt. of India. Johnsingh, A.J.T. (1992). Prey selection in three large sympatric carnivores in Bandipur. Mammalia, 56, 517-526. https://doi. org/10.1515/mamm.1992.56.4.517
- Johnsingh, A.J.T., and Negi, A.S. (2003). Status of tiger and leopard in Rajaji-Corbett Conservation Unit, northern India. Biological Conservation, 111, 385-393. https://doi.org/10.1016/S0006-3207(02)00307-5
- Kalle, R., Ramesh, T., Qureshi, Q., and Sankar, K. (2011). Density of tiger and leopard in a tropical deciduous forest of Mudumalai Tiger Reserve, southern India, as estimated using photographic capture-recapture sampling. Acta Theriologica, 56, 335-342. https://doi.org/10.1007/s13364-011-0038-9
- Karanth, K. (2003). Tiger ecology and conservation in the Indian subcontinent. Journal of the Bombay Natural History Society, 100, 169-189.
- Karanth, K., Nichols, J., Seidenstricker, J., Dinerstein, E., Smith, J., McDougal, C., et al. (2003). Science deficiency in conservation practice: the monitoring of tiger populations in India. Animal Conservation Forum, 6, 141-146. https://doi.org/10.1017/S1367943003003184
- Karanth, K.U. (1995). Estimating tiger Panthera tigris populations from camera-trap data using capture-recapture models. Biological Conservation, 71, 333-338. https://doi.org/10.1016/0006-3207(94)00057-W
- Karanth, K.U., and Nichols, J.D. (1998). Estimation of tiger densities in India using photographic captures and recaptures. Ecology, 79, 2852-2862. https://doi.org/10.1890/0012-9658(1998)079%5b2852:EOTDII%5d2.0.CO;2
- Karanth, K.U., and Nichols, J.D. (2002). Monitoring Tigers and Their Prey: A Manual for Wildlife Researchers, Managers and Conservationists in Tropical Asia. Centre for Wildlife Studies.
- Karanth, K.U., and Sunquist, M.E. (1995). Prey selection by tiger, leopard and dhole in tropical forests. Journal of Animal Ecology, 64, 439-450. https://doi.org/10.2307/5647
- Karanth, K.U., Chundawat, R.S., Nichols, J.D., and Kumar, N.S. (2004a). Estimation of tiger densities in the tropical dry forests of Panna, Central India, using photographic capture-recapture sampling. Animal Conservation, 7, 285-290. https://doi.org/10.1017/S1367943004001477
- Karanth, K.U., Nichols, J.D., Kumar, N.S., Link, W.A., and Hines, J.E. (2004b). Tigers and their prey: predicting carnivore densities from prey abundance. Proceedings of the National Academy of Sciences of the United States of America, 101, 4854-4858. https://doi.org/10.1073/pnas.0306210101
- Kitchener, A. (1991). The Natural History of the Wild Cats. Comstock Pub.
- Majumder, A. (2011). Prey selection, food habits and population structure of sympatric carnivores: Tiger panthera tigris tigris (L.), Leopard Panthera pardus (L.) and Dhole Cuon alpinus (Pallas) in Pench Tiger Reserve, Madhya Pradesh (India). (doctoral dissertation). Saurastra University, Rajkot.
- Majumder, A., Basu, S., Sankar, K., Qureshi, Q., Jhala, Y.V., Nigam, P., et al. (2012). Home ranges of the radio-collared Bengal tigers (Panthera tigris tigris L.) in Pench Tiger Reserve, Madhya Pradesh, Central India. Wildlife Biology in Practice, 8, 36-49.
- Mondal, K., Gupta, S., Bhattacharjee, S., Qureshi, Q., and Sankar, K. (2012). Response of leopards to re-introduced tigers in Sariska Tiger Reserve, Western India. International Journal of Biodiversity and Conservation, 4, 228-236. https://doi.org/10.5897/IJBC12.014
- Nowak, R.M. (1999). Walker's Mammals of the World, 6th ed. Johns Hopkins University Press.
- Nowell, K., and Jackson, P. (1996). Wild Cats: Status Survey and Conservation Action Plan. IUCN.
- Otis, D.L., Burnham, K.P., White, G.C., and Anderson, D.R. (1978). Statistical inference from capture data on closed animal populations. Wildlife Monographs, 62, 3-135.
- Panda, D., Mohanty, S., Allen, M.L., Dheer, A., Sharma, A., Pandey, P., et al. (2023).Competitive interactions with dominant carnivores affect carrion acquisition of striped hyena in a semi-arid landscape of Rajasthan, India. Mammal Research, 68, 129-141. https://doi.org/10.1007/s13364-022-00663-1
- Panda, D., Mohanty, S., Suryan, T., Pandey, P., Lee, H., and Singh, R. (2022). High striped hyena density suggests coexistence with humans in an agricultural landscape, Rajasthan. PloS One, 17, e0266832. https://doi.org/10.1371/journal.pone.0266832
- Panwar, H.S. (1979). A note on tiger census technique based on pugmark tracings. Tigerpaper, 6, 16-18.
- Ramesh, T., Snehalatha, V., Sankar, K., and Qureshi, Q. (2009). Food habits and prey selection of tiger and leopard in Mudumalai Tiger Reserve, Tamil Nadu, India. Journal of Scientific Transactions in Environment and Technovation, 2, 170-181. https://doi.org/10.20894/STET.116.002.003.010
- Rexstad, E., and Burnham, K.P. (1991). User's Guide for Interactive Program CAPTURE. Cooperative Fish and Wildlife Research Unit.
- Rodgers, W.A., and Panwar, S.H. (1988). Biogeographical Classification of India. New Forest.
- Sehgal, J.J., Kumar, D., Kalsi, R.S., Allen, M.L., and Singh, R. (2022). Spatio-temporal overlap of leopard and prey species in the foothills of Shiwalik, Himalaya. European Journal of Wildlife Research, 68, 18. https://doi.org/10.1007/s10344-022-01568-9
- Seidensticker, J. (1976). On the ecological separation between tigers and leopards. Biotropica, 8, 225-234. https://doi.org/10.2307/2989714
- Seidensticker, J., and Sunquist, M.E. (1990). Leopards living at the edge of the Royal Chitwan National Park, Nepal. In J.C. Daniel, and J.S., Serrao (Eds.), Conservation in Developing Countries: Problems and Prospects (pp. 415-423). Oxford University Press.
- Singh, R., Krausman, P.R., Goyal, S.P., and Chauhan, N.S. (2015). Factors contributing to tiger losses in Ranthambhore Tiger Reserve, India. Wildlife Society Bulletin, 39, 670-673. https://doi.org/10.1002/wsb.561
- Singh, R., Pandey, P., Qureshi, Q., Sankar, K., Krausman, P.R., and Goyal, S.P. (2020). Acquisition of vacated home ranges by tigers. Current Science, 119, 1549-1554. https://doi.org/10.18520/cs/v119/i9/1549-1554
- Singh, R., Pandey, P., Qureshi, Q., Sankar, K., Krausman, P.R., and Goyal, S.P. (2021). Philopatric and natal dispersal of tigers in a semi-arid habitat, western India. Journal of Arid Environments, 184, 104320. https://doi.org/10.1016/j.jaridenv.2020.104320
- Singh, R., Pandey, P., Qureshi, Q., Sankar, K., Krausman, P.R., and Goyal, S.P. (2022). Temporal variation in tiger population in a semi-arid habitat in India. Proceedings of the National Institute of Ecology of the Republic of Korea, 3, 154-164. https://doi.org/10.22920/PNIE.2022.3.3.154
- Singh, R., Qureshi, Q., Sankar, K., Krausman, P.R., and Goyal, S.P. (2013). Use of camera traps to determine dispersal of tigers in semi-arid landscape, western India. Journal of Arid Environments, 98, 105-108. https://doi.org/10.1016/j.jaridenv.2013.08.005
- Singh, R., Qureshi, Q., Sankar, K., Krausman, P.R., and Goyal, S.P. (2014). Evaluating heterogeneity of sex-specific capture probability and precision in camera-trap population estimates of tigers. Wildlife Society Bulletin, 38; 791-796. https://doi.org/10.1002/wsb.471
- Thapa, K., Shrestha, R., Karki, J., Thapa, G.J., Subedi, N., Pradhan, N.M.B., et al. (2014). Leopard Panthera pardus fusca density in the seasonally dry, subtropical forest in the Bhabhar of Terai Arc, Nepal. Advances in Ecology, 2014, 286949. https://doi.org/10.1155/2014/286949
- Thapa, T.B. (2011). Habitat suitability evaluation for leopard (Panthera pardus) using remote sensing and GIS in and around Chitwan National Park, Nepal. (Unpublished doctoral dissertation or master's thesis). Saurashtra University, Rajkot.
- Wilson, G.J., and Delahay, R.J. (2001). A review of methods to estimate the abundance of terrestrial carnivores using field signs and observation. Wildlife Research, 28, 151-164. https://doi.org/10.1071/WR00033
- WPSI. (2006). Skinning the cat: crime and politics of the big cat skin trade. Retrieved February 21, 2021 from http://www.indiaenvironmentportal.org.in/files/Skinning%20the%20Cat.pdf.
- Zimmermann, F., Breitenmoser-Wursten, C., Molinari-Jobin, A., and Breitenmoser, U. (2013). Optimizing the size of the area surveyed for monitoring a Eurasian lynx (Lynx lynx) population in the Swiss Alps by means of photographic capturerecapture. Integrative Zoology, 8, 232-243. https://doi. org/10.1111/1749-4877.12017