• Journal of Ecology and Environment
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• 제41권1호
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• pp.1-8
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• 2017

Background: Though the biomass of floral vegetation in understory plant communities in a forested ecosystem only accounts for less than 1% of the total biomass of a forest, they contain most of the floral resources of a forest. The diversity of understory honey plants determines visitation rate of pollinators such as honey bee (Apis mellifera) as they provide rich food resources. Since the flower visitation and foraging activity of pollinators lead to the provision of pollination service, it also means the enhancement of plant-pollinator relationship. Therefore, an appropriate management scheme for understory vegetation is essential in order to conserve pollinator population that is decreasing due to habitat destruction and disease infection. This research examined the diversity of understory honey plant and studied how it is related to environmental variables such as (1) canopy density, (2) horizontal heterogeneity of canopy surface height, (3) slope gradient, and (4) distance from roads. Vegetation survey data of 39 plots of mixed forests in Chuncheon, Korea, were used, and possible management practices for understory vegetation were suggested. Results: This study found that 113 species among 141 species of honey plant of the forests were classified as understory vegetation. Also, the understory honey plant diversity is significantly positively correlated with distance from the nearest road and horizontal heterogeneity of canopy surface height and negatively correlated with canopy density. Conclusions: The diversity of understory honey plant vegetation is correlated to vegetation structure and human impact. In order to enhance the diversity of understory honey plant, management of density and height of canopy is necessary. This study suggests that improved diversity of canopy cover through thinning of overstory vegetation can increase the diversity of understory honey plant species.

• Journal of Ecology and Environment
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• 제32권2호
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• pp.87-96
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• 2009

Pinus thunbergii plantations in Pohang-si, Gyeongsangbuk-do, Korea, are of low ecological quality, with arrested succession and a high proportion of ruderal species. To improve the quality of the habitat, we created canopy gaps ($\sim42\;m^2$) and monitored changes in abiotic (light availability, canopy openness) and biotic (survival and growth of seedlings and understory communities) variables in 2007 and 2008 in plots that had received one of five types of treatment: cutting of canopy trees and removal of the understory (CU), cutting of canopy trees only (C), girdling of canopy trees and removal of the understory (GU), girdling of canopy trees (G) or control. Each treatment was applied to three replicate plots. Abiotic variables did not significantly differ among treatments. Survival rates of target species were slightly lower in the CU, G and control conditions. Based on logistic regression analysis, the only significant growth factor affecting survival was height growth. Positive effects of seedling height and leaf area growth on survival were also detected, but did not reach statistical significance. In treatment G, gradual improvement of overstory conditions and mitigation of competition by limitation of disturbance to the understory community were likely to have promoted seedling growth. There were no significant effects of gap treatments on changes in species abundance (cover and richness) and composition of understory between the study years. This result implies that the small gaps created in our study may be below the threshold size to affect understory growth. However, the results of this study are based on a short-term investigation of only two years. Long-term research is strongly recommended to clarify the effects of gap treatment on plant communities in afforested areas.

• The Korean Journal of Ecology
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• 제12권4호
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• pp.245-256
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• 1989

Nitrogen cycling was in vestigated in Piuns koraiensis plantations with different ages, 1, 2, 3, 6, 9 and 11 years, which were reforested after clear-cutting. Annual N input by bulk precipitation was 10kg., and output by runoff decreased as the plantation aged, especially in-N. The standing N content of the whole vegetation increased approximately 5 times through 11 years. Understory surpassed P. koraiensis plants in the distribution of standing N content for the initial 9 years, but reversed thereafter. Annual N uptake of P. koraiensis plants increased greatly through 11 years, but that of understory increased somewhat until 9 years and decreased thereafter. The maximum N uptake of the whole vegetation was made in the 9-yr-old plantation. In the 1-yr-old one, 59% of the maximum was already absorbed by understory which mainly consisted to herbs. The recycling coefficient, ratio of annual return to rptake, of the whole vegetation decreased as the plantation aged and the value of understory was greater than that of P koraiensis plants. On the contrary, the N use efficiency, ratio of the net primary production to N uptake, of the whole vegetation increased as the plantation aged and the value of understory was less than that of P. koraiensis plants. Consequently, it is emphasized that understory played an important role in such plantation reforesred after clear-cutting for the initial 9 years.

• 한국산림과학회지
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• 제96권2호
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• pp.151-159
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• 2007

This study was conducted to investigate the effects of thinning on changes in stand characteristics and understory vegetation in a silvopasture practiced Japanese larch plantation in the Research Forest of Kangwon National University, Korea. Three different thinning intensities (64%, 35%, and control) were applied. Before and after thinning, the understory plant species increased its number from 48 (7 tree species, 7 shrubs species, 28 herbaceous species, and 6 woody climbers) to 100 (11 tree species, 15 shrub species, 67 herbaceous species, and 7 woody climbers). Thinning made plants invade easily on the forest floor, and plot A (325 stems/ha) had much higher number of undersory species than those Of plot B (575 stems/ha) and control plot (1,150 stems/ha). In three years after thinning, understory aboveground biomass (kg/ha) of herbs were 523 for control, 1,230 for plot B, and 1,288 for plot A. The canopy coverage had remarkable influence on the understory biomass production, resulting in relatively small amount of herbage production on control plot. The differences were statistically significant between thinned plots and unthinned plot, but there were no significant differences among the thinned plots (p<0.05).

• Animal cells and systems
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• 제3권2호
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• pp.143-147
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• 1999

Annual production and redistribution of leaf litter were compared among three distinct understory patches in a temperate hardwood forest dominated by Quercus mongolica, Kalopanax pictus, Acer pseudo-sieboldianum, and Carpinus cordata. Two patches were located on a southwest-facing slope: one with an understory dominated by herbaceous plants (Patch S), and the other covered with evergreen dwarf bamboo, Sasa borealis (patch SS). The third patch was on the opposite slope with an understory dominated by herbaceous plants (Patch N). Annual leaf litterfall was averaged 330 g m$^{-2} yr$^{-2}$ in the three patches from 1994 to 1998. From mid-September 1996 to mid-September 1997, net transport of leaf litter over patch bound-aries was 1,824g m$^{-1}$ from Patch S to SS, 1,465g m$^{-1}$ from Patch S to N, and 886 g m$^{-1}$ from Patch SS to N. The amounts moving downslope out of Patch S, SS, and N were 2,548, 471, and 588g m$^{-1}$, respectively. When a mass balance approach was employed for the data of leaf litter transport, the results were relatively consistent with 216, 631, and 724g m$^{-2}$ of leaf litter stores in Patch S, SS, and N, respectively, in April 1997. This study suggests that leaf litter redistribution is largely regulated by aspect and understory type and exerts a significant effect on carbon processes in the forest ecosystem.

• 한국산림과학회지
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• 제90권3호
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• pp.236-241
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• 2001

본 연구는 산림 하층의 환경특성과 설치류 서식밀도를 조사하여 산림생태계 내에서 소형 설치류와 산림 하층의 서식환경과의 관계를 파악하고자 1997년 7월부터 1998년 8월 사이에 지리산국립공원의 6개 지역에서 각각 해발고도 500m, 800m, 1,100m에 설정한 총 18개의 조사 지역에서 실시하였다. 해발고도가 높아짐에 따라 대부분의 지역에서 하층식생의 피도량과 유기물층의 깊이가 증가하는 것으로 나타났다. 소형 설치류의 포획결과 등줄쥐, 흰넓적다리붉은쥐, 대륙밭쥐 등 3종 77개체가 포획되었는데 각 지역별 포획 개체수는 하층식생의 피도량과 유기물층의 발달 정도와 밀접한 관련이 있는 것으로 나타났다. 토양내 하층식생의 발달로 인한 피도량의 증가와 유기물층의 발달은 소형 설치류에게 있어서 알맞은 서식환경을 제공해 주는 것으로 판단된다.

• 한국산림과학회지
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• 제103권1호
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• pp.23-29
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• 2014

천이 모형과 같이 임분 교란 후 수관 닫힘 단계까지의 다양한 생태적 특성 변화에 관한 많은 연구가 있으나 우리나라는 다양한 연구가 수행되고 있지 않다. 본 연구는 강원도 남부 소나무림 개벌 후 임분 구조, 환경 및 하층식생 풍부성의 경시적 변화 추이를 발달 단계별(1yr, 3yr, 10yr 및 16yr)로 분석하였다. 임분구조 분석 결과, 벌채 후, 평균 줄기 밀도는 16yr에 $5,714{\pm}645$ stems/ha 까지 점차 증가하였고, 평균 기저면적 역시 10yr에 $5.5{\pm}0.7m^2/ha$, 그리고 이후 $16yr(10.0{\pm}1.6m^2/ha)$)까지 거의 두 배로 증가함을 알 수 있었다. 숲 바닥의 나무 잔해 및 토양 노출도는 16yr의 11% 및 3yr의 10.3%에서 각각 최고점을 나타낸 후 감소하는 경향이었다. 하층식생 피도는 16년 동안의 임분구조 발달에 따라 모든 생육형에서 모두 감소하는 경향이었으나, 광엽 초본의 종풍부도는 반대로 증가하는 경향을 나타내었다. 특히, 임분의 상층식생 발달은 하층 식생 속성을 억누르기 보다는 필터 역할과 같이 선택적 영향을 미치는 것으로 생각되었다. 본 연구와 더불어 보다 세밀한 생태 속성 변화를 관찰하기 위해서는 임분의 초기 발달 패턴과 과정에 대한 연속 모니터링이 중요하다.

• 한국농림기상학회지
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• 제11권4호
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• pp.233-246
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• 2009

증발산(ET)을 증발(E)과 증산(T)으로 배분하는 것은 물 순환과 에너지, 물 및 탄소 순환의 연결고리를 이해하는 데에 매우 중요하다. 산림 군락의 총 증발산은 상부 및 하부 군락의 증산과 토양 및 차단 강수로부터의 증발로 구성된다. 이들의 상대적 기여도를 정량화하기 위해, 에디 공분산 방법을 사용하여 2008년 6월 1일부터 2009년 5월 31일까지 광릉 활엽수림과 침엽수림의 마루에서의 증발산을 관측하였다. 스펙트럼 분석에 따르면, 초음파 풍향 풍속계와 수증기 농도 측정기 사이의 거리에 의한 증발산 과소 평가는 군락 하부에서 연간 증발산량의 약 10%정도였다. 하부군락의 연간 증발산은 활엽수림과 침엽수림에서 각각 59mm와 43mm로, 총 증발산량의 16%와 7%를 차지하였다. 전반적으로 식생면적지수가 최대인 여름 기간을 제외하고는 하부군락의 증발산은 총 증발산에서 무시할 수 없는 부분을 차지하였다. 두 산림지역의 비결합 모수($\Omega$)는 약 0.15이었으며, 이는 하부군락의 증발산이 주로 포차나 토양 수분에 의해서 조절됨을 보여준다. 두 산림의 하부 군락의 증발산의 차이는 군락 하부의 환경 조건의 차이에 기인하며, 특히 습도와 토양 수분의 차이로 인해 발생하였다. 광릉 하부 군락의 증발산 기여도가 작지 않다는 사실은 이러한 산림의 물질과 에너지 플럭스를 해석하고 모델링하기 위해서는 이원 또는 다층 모형이 필요함을 시사한다.

• 한국산림과학회지
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• 제112권3호
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• pp.363-373
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• 2023

산림 바이오매스 조사는 탄소흡수원으로서의 산림을 평가하고 관리하기 위해 주기적으로 수행된다. 원격탐사의 한 종류인 라이다는 적은 노동력으로 객관적인 산림 구조 정보를 획득할 수 있어, 최근 라이다(LiDAR, Light Detection and Ranging)를 이용한 산림 조사가 주목받고 있다. 본 연구에서는 임분 상하층 바이오매스 추정에 백팩형 라이다(Backpack Laser Scanning, BPLS)와 드론 라이다(Unmanned Aerial Vehicle Laser Scanning, UAV-LS)를 이용하는 방법을 제시하고 그 정확도를 평가하였다. 상층의 경우 BPLS와 UAV-LS의 흉고직경과 수고 추정 정확도를 분석하였고, 하층의 경우 BPLS 데이터에서 추출한 수직구조 변수 중 최상의 변수 조합으로 하층 바이오매스를 추정하는 다중회귀모델을 개발하였다. 그 결과, BPLS는 흉고직경을 높은 정확도로 추정하였지만(R² =0.92) 수고는 과소 추정하였다(R² =0.63, Bias=-5.56 m). UAV-LS는 BPLS보다 더 높은 수고 추정 정확도를 보였다(R² =0.91). 하층의 경우 점들의 평균 높이와 라이다 데이터를 같은 높이를 가진 10개의 층으로 나누었을 때 아래에서 네 번째 층의 점 밀도를 의미하는 변수가 선택되어 모델이 개발되었으며, 교차검증 결과 결정계수 값은 0.68로 나타났다. 본 연구의 결과는 BPLS와 UAV-LS를 이용한 임분의 상하층 바이오매스 조사 방법이 기존의 조사 방식을 효과적으로 대체할 수 있음을 시사한다.

• 한국산림과학회지
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• 제96권5호
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• pp.520-527
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• 2007

Biomass and species diversity of understory vegetation after fertilization were studied for 28-year-old Quercus acutissima plantation (MQA), 29-year-old Pinus densiflora plantation (MPD), 8-year-old Betula platyphylla var. japonica plantation after coal reclamation (YBP), and 4-year-old Pinus densiflora plantation after forest fire (YPD) in central Korea. Nitrogen + phosphorus + potassium (6:4:1) fertilizer was applied for 3 years from 2004. Thereafter photosynthetically active radiation (PAR) and understory species richness and diversity were measured in late July-early August 2006. PAR (${\mu}mol\;m^{-2}S^{-1}$) was higher at the fertilization treatment (100.9) than at the control (67.0) for MQA while was lower at the fertilization treatment (156.5) than at the control (268.7) for MPD. Total understory biomass (t $ha^{-1}$) was lower at the fertilization plot (1.8) than at the control plot (3.0) for YPD, however, there were no differences in biomass between fertilization and control plots for MQA, MPD and YBP. Total species number of understory vegetation was higher for fertilization than for control at MPD (47 vs. 45) and YPD (21 vs. 13), and was higher for mature plantations (33 vs. 37 for MQA and 47 vs. 45 for MPD) than for young plantations (16 vs. 16 for YBP and 21 vs. 13 for YPD). Species richness and diversity were higher at the fertilization treatment than at the control for MQA, YBP, and YPD while were lower at the fertilization treatment than at the control for MPD, however, the differences were not statistically significant. Our results indicate that there were no consistent patterns in light conditions, biomass and species richness and diversity of understory vegetation following fertilization. More detailed long-term studies with different fertilizer applications would be necessary to conclude the influence of fertilization on understory vegetation in the region.