• Title/Summary/Keyword: Morphological evolution

Search Result 396, Processing Time 0.028 seconds

한국산 빗살거미불가사리속(빗살거미불가사리과, 거미불가사리아강)의 3 종에 대한 분류학적 재검토

  • 유재원;홍재상;박흥식
    • Animal Systematics, Evolution and Diversity
    • /
    • v.11 no.4
    • /
    • pp.417-434
    • /
    • 1995
  • Three species (0. kinbergi, 0. sarsi and 0 . sarsi vadicola) of the genus Ophiura (Echinodermacta: Ophiuroidea) were sampled from the various localities of Korean waters (Kyonggi Bay in March and September, 1989 and February, 1994; Yellow Sea in September and October, 1992; Southern Sea and Korean Strait in May, 1992; and eastern coasts adjacent to Kangnung in April, July, October, 1993 and January, 1994). Results of the examination of 250 Operational Taxonomical Units (OTUs) are presented based on the 20 morphometric variables to evaluate their taxonomic characters and positions. In cluster analysis, 250 OTUs were divided into 3 phenons (0. kinbergi, 0 . sarsi and 0 . sarsi vadicola) at the Euclidean distance levels of 6.84 and 2 phenons (a phenon composed of 0 . sarsi and 0 . sarsi vadicola and the other of 0. kinbergi) at 7.50. Stepwise discriminant analysis was used in order to produce a good discrimination model and 13 morphological characters (the total number of comb papillae, the number of primary comb papillae and shape of comb papillae (2). etc.) were extracted. The results of canonical discriminant analysis illustrated clear distinction among 3 phenons by the distance of 8.26 between 0 . sarsi and 0. sarsi vadicola, 24.24 between 0 . kinbergi and 0. sarsi vadicola and 21.63 between 0 . kinbergi and 0 . sarsi.

  • PDF

Phylogeny of the Polygonatum odoratum Complex Inferred from Multiple cpDNA and Nuclear RNApol2_i23 Sequence Data (Ruscaceae) (엽록체 DNA 및 핵 DNA RNApol2_i23에 근거한 둥굴레복합체 (Ruscaceae)의 계통 연구)

  • Park, Jeong-Mi;Chung, Kyong-Sook;Oh, Byoung-Un;Jang, Chang-gee
    • Korean Journal of Plant Taxonomy
    • /
    • v.41 no.4
    • /
    • pp.353-360
    • /
    • 2011
  • The sequence data of the plastid DNA (trnL-F IGS, trnL intron, and trnH-psbA) and nuclear DNA (RNApol2_i23) markers were utilized to study phylogenetic relationships among the taxa in the Polygonatum odoratum complex (Ruscaceae). European P. odoratum individuals form a clade with a high bootstrap value, which is a sister to the clade of Korean P. odoratum var. odoratum, P. odoratum var. pluriflorum and P. robustum. The formation of the clade with P. odoratum var. odoratum, P. robustum, and one accession of P. odoratum var. pluriflorum indicates geological speciation in isolated populations in the islands following dispersal events from the mainlands. All data sets form two major clades, which are congruent with the subgroups divided by the basic chromosome numbers (x = 9 and x = 10). Although it is not easy to test the hypothesis of the decrease in the basic chromosome number due to scatter taxon sampling in this study, the molecular data strongly suggested that aneuploidy plays an important role in lineage diversification in the genus Polygonatum. The cytological data was not strongly supported by the cpDNA sequences. Further investigations of the cytological, morphological, and geographical characteristics with comprehensive sampling are desired to understand the evolution and lineage diversification in the genus.

Impacts of wave and tidal forcing on 3D nearshore processes on natural beaches. Part I: Flow and turbulence fields

  • Bakhtyar, R.;Dastgheib, A.;Roelvink, D.;Barry, D.A.
    • Ocean Systems Engineering
    • /
    • v.6 no.1
    • /
    • pp.23-60
    • /
    • 2016
  • The major objective of this study was to develop further understanding of 3D nearshore hydrodynamics under a variety of wave and tidal forcing conditions. The main tool used was a comprehensive 3D numerical model - combining the flow module of Delft3D with the WAVE solver of XBeach - of nearshore hydro- and morphodynamics that can simulate flow, sediment transport, and morphological evolution. Surf-swash zone hydrodynamics were modeled using the 3D Navier-Stokes equations, combined with various turbulence models (${\kappa}-{\varepsilon}$, ${\kappa}-L$, ATM and H-LES). Sediment transport and resulting foreshore profile changes were approximated using different sediment transport relations that consider both bed- and suspended-load transport of non-cohesive sediments. The numerical set-up was tested against field data, with good agreement found. Different numerical experiments under a range of bed characteristics and incident wave and tidal conditions were run to test the model's capability to reproduce 3D flow, wave propagation, sediment transport and morphodynamics in the nearshore at the field scale. The results were interpreted according to existing understanding of surf and swash zone processes. Our numerical experiments confirm that the angle between the crest line of the approaching wave and the shoreline defines the direction and strength of the longshore current, while the longshore current velocity varies across the nearshore zone. The model simulates the undertow, hydraulic cell and rip-current patterns generated by radiation stresses and longshore variability in wave heights. Numerical results show that a non-uniform seabed is crucial for generation of rip currents in the nearshore (when bed slope is uniform, rips are not generated). Increasing the wave height increases the peaks of eddy viscosity and TKE (turbulent kinetic energy), while increasing the tidal amplitude reduces these peaks. Wave and tide interaction has most striking effects on the foreshore profile with the formation of the intertidal bar. High values of eddy viscosity, TKE and wave set-up are spread offshore for coarser grain sizes. Beach profile steepness modifies the nearshore circulation pattern, significantly enhancing the vertical component of the flow. The local recirculation within the longshore current in the inshore region causes a transient offshore shift and strengthening of the longshore current. Overall, the analysis shows that, with reasonable hypotheses, it is possible to simulate the nearshore hydrodynamics subjected to oceanic forcing, consistent with existing understanding of this area. Part II of this work presents 3D nearshore morphodynamics induced by the tides and waves.

Wind-sand coupling movement induced by strong typhoon and its influences on aerodynamic force distribution of the wind turbine

  • Ke, Shitang;Dong, Yifan;Zhu, Rongkuan;Wang, Tongguang
    • Wind and Structures
    • /
    • v.30 no.4
    • /
    • pp.433-450
    • /
    • 2020
  • The strong turbulence characteristic of typhoon not only will significantly change flow field characteristics surrounding the large-scale wind turbine and aerodynamic force distribution on surface, but also may cause morphological evolution of coast dune and thereby form sand storms. A 5MW horizontal-axis wind turbine in a wind power plant of southeastern coastal areas in China was chosen to investigate the distribution law of additional loads caused by wind-sand coupling movement of coast dune at landing of strong typhoons. Firstly, a mesoscale Weather Research and Forecasting (WRF) mode was introduced in for high spatial resolution simulation of typhoon "Megi". Wind speed profile on the boundary layer of typhoon was gained through fitting based on nonlinear least squares and then it was integrated into the user-defined function (UDF) as an entry condition of small-scaled CFD numerical simulation. On this basis, a synchronous iterative modeling of wind field and sand particle combination was carried out by using a continuous phase and discrete phase. Influencing laws of typhoon and normal wind on moving characteristics of sand particles, equivalent pressure distribution mode of structural surface and characteristics of lift resistance coefficient were compared. Results demonstrated that: Compared with normal wind, mesoscale typhoon intensifies the 3D aerodynamic distribution mode on structural surface of wind turbine significantly. Different from wind loads, sand loads mainly impact on 30° ranges at two sides of the lower windward region on the tower. The ratio between sand loads and wind load reaches 3.937% and the maximum sand pressure coefficient is 0.09. The coupling impact effect of strong typhoon and large sand particles is more significant, in which the resistance coefficient of tower is increased by 9.80% to the maximum extent. The maximum resistance coefficient in typhoon field is 13.79% higher than that in the normal wind field.

Systematic Studies on Korean Rodents : VIII. Analyses of Morphometric Characters, Chromosomal Karyotype, and Mitochondrial DNA Restriction Fragments in Siberian Chipmunks from Korea (Tamias sibiricus barberi Johnson and Jones), with the Comparison of Morphometric Characters of Siberian Chipmunks from Manchuria (Tamias sibiricus orientalis Bonhote) (한국산 설치류의 계통분류학적 연구: 8. 다람쥐(Tamias sibiricus barberi Johnson and Jones)의 형태적 형질, 염색체 핵형 및 미토콘드리아 DNA절단 단편의 분석과 만주다람쥐(Tamias sibiricus orientalis Bonhote)와의 형태적 형질의 비교)

  • Hung Sun Koh
    • Animal Systematics, Evolution and Diversity
    • /
    • v.10 no.2
    • /
    • pp.231-243
    • /
    • 1994
  • Samples of Siberian chipmunks (Tamias sibiricus borberi ) from Korea were analyzed by multivariate, air-drying , and blot-hybridization methods in order to determine the degree of variations of morphological characters, chromosomal karyotype, and mtDNA restriction fragment patterns, respectively, Moreover, morphometric characters of samples of Siberian chipmunks from Manchuria(T.sibiricus orientalis) were also analyzed in order to clarify the taxonomic status of the subspecies barberi. Siberian chipmunks from six localities in Korea were in morphometric characters more or less similar with one another and Siberian chipmunks from Manchuria were not so distinct enough to from a subgroup. Siberian chipmunks from Four localities in Korea were identical in their karyotypes (2n=38) : samples from three localities in Korea were nore or less similar in their mtDNA fragment patterns. Siberian chipmunks from Korea appeared to be a omogeneous population and as noted by Corbet (1978) subspecies barberi is the synonym of subspecies orientalis. The scientific name of Siberian chipmunks from Korea and Manchuria is T. sibiricus orientalis, although systematic studies with the samples from North korea, Manchuria and China will be necessary for their taxonomic reconsideration.

  • PDF

On the Debates of Arthropod Phylogeny (절지동물 계통에 관한 논쟁)

  • 황의욱
    • Animal Systematics, Evolution and Diversity
    • /
    • v.18 no.1
    • /
    • pp.165-179
    • /
    • 2002
  • In spite of dramatic change of environmental condition since Cambrian big-bang (explosion occurred ca.540 mya, the phylum Arthropoda retains a great diversity, and it is estimated approximately that 1-10 million arthropod species are extant on the earth. Except for an extinct arthopod subphylum Trilobita, extant arthropods could be divided into five subphyla: Hexapoda, Crustacea, Myriapoda, Chelicerata, and Pycnosonida. During the last century, systematists have disputed about interrealtionships among Arthropoda and its relatives (Onychophora, Tardigrada, and Pentastomida), arthropod phylogenetic position within protostome animals, monophyly or polyphyly of the phylum Arthropods, and interrelationships among five arthropod subgroups (subphyla) etc. Recently, new animal phylogeny was reported that protostomes could be clustered into two groups, Lophotrochozoa and Ecdysozoa, and molting animals such as Nematoda and Arthropoda were included within the Ecdysozoa. On the basis of the new animal phylogeny, first of all, I would mention phylogenetic positions and relationships of Arthropods and its relatives to introduce controversies of arthropod phylogeny in phylum level of animals. After that, I focused mainly on the controveries related to arthropod monophyly and phylogenetic relationships among four major arthropod groups except Pycnogonida. In this work, Pycnogonida which is a relatively small group and one of the five arthropod subphyla was not handled significantly although there are some controversies if it is a sister taxon of chelicerates or the most primitive arthropod group (namely, a sister of four remains arthropod groups).

Early-type Dwarf Galaxies in the Virgo Cluster: An Ultraviolet Perspective

  • Kim, Suk;Rey, Soo-Chang;Sung, Eon-Chang;Lisker, Thorsten;Jerjen, Helmut;Lee, Youngdae;Chung, Jiwon;Pak, Mina
    • The Bulletin of The Korean Astronomical Society
    • /
    • v.37 no.2
    • /
    • pp.81-81
    • /
    • 2012
  • Since the ultraviolet (UV) flux of an integrated population is a good tracer of recent star formation activities, UV observations provide an important constraint on star formation history (SFH) in galaxies. We present UV color-magnitude relations (CMRs) of early-type dwarf galaxies in the Virgo cluster, based on Galaxy Evolution Explorer (GALEX) UV data and the Extended Virgo Cluster Catalog (EVCC, Kim, S. in prep.). The EVCC covers an area 5.4 times larger (750 deg2) than the footprint of the classical Virgo cluster catalog by Binggeli and collaborators. We secure 1304 galaxies as members of the Virgo cluster and 526 galaxies of them are new objects not contained in the VCC. Morphological classification of galaxies in the EVCC is based on the optical image ("Primary Classification") and spectral feature ("Secondary Classification") of the SDSS data. We find that dwarf lenticular galaxies (dS0s) show a surprisingly distinct and tight locus separated from that of ordinary dwarf elliptical galaxies (dEs), which is not clearly seen in previous CMRs. The dS0s in UV CMRs follow a steeper sequence than dEs and show bluer UV-optical color at a given magnitude. Most early type dwarf galaxies with blue UV colors (FUV-r < 6 and NUV-r < 4) are identified as those showing spectroscopic hints of recent or ongoing star formation activities. We explore the observed CMRs with population models of a luminosity-dependent delayed exponential star formation history. The observed CMR of dS0s is well matched with models with relatively long delayed star formation. Our results suggest that dS0s are most likely transitional objects at the stage of subsequent transformation of late-type progenitors to ordinary red dEs in the cluster environment. In any case, UV photometry provides a powerful tool to disentangle the diverse subpopulations of early-type dwarf galaxies and uncover their evolutionary histories.

  • PDF

Redescription of Two Terrestrial Hypotrichous Ciliates from Jindo Island, Korea (진도의 육상 하모섬모충류 2미기록 종의 재기재)

  • Kwon, Choon-Bong;Shin, Mann-Kyoon
    • Animal Systematics, Evolution and Diversity
    • /
    • no.nspc5
    • /
    • pp.71-79
    • /
    • 2005
  • Two hypotrichous ciliates inhabiting the representative terrestrial habitats of Jindo Island, Holostichides chardezi Foissner, 1987 and Rigidocortex octonucleatus (Foissner, 1988) are found from Jindo Island. These two species are newly recorded from Korea. The description was based on the observation of living specimens, protargol impregnated specimens and biometric analysis. The morphological variations among the populations of these species were investigated with morphometry. The diagnostic characteristics of these species are as follows: Holostichides chardezi : elongate body flexible and twisted, yellowish subpellicular granules scattered entire cortex, adoral zone of membranelles covering about $28\%$ of body length, left and right rows of mid-ventral cirri paired, left row ended at buccal vertex and right row extending to center of body, no transverse cirri, both marginal rows of cirri not confluent at posterior end, 28-42 macronuclei and 1-5 micronuclei scattered entire body. 3 caudal cirri and 4 dorsal kineties. Rigidocortex octonucleatues: elongate-oval body rigid and not contractile, yellowish cytoplasmic crystals and fat globules densely packed, adoral zone of membranelles covering about $40\%$ of body length, 5 transverse cirri, both marginal rows of cirri not confluent at posterior end, 8 macronuclei longitudinally arranged in median of cell, 3 caudal cirri and 7-10 fragmented dorsal kineties.

The Summer Growth and the Ratios of the Breadth to the Length of the Carapace of the Crab, Macrophthalmus (Mareotis) japonicus De Hann (강화산 칠게의 여름철 성장 및 갑폭과 중장의 비)

  • Kim, Hoon-Soo;Kwak, Hi-Sang;Kwon, Do-Heon
    • Animal Systematics, Evolution and Diversity
    • /
    • no.nspc2
    • /
    • pp.233-241
    • /
    • 1988
  • The crab, Macrophthalmus (Mareotis ) japonicus , collected in 1983 from CHojijin tidal flat, southeastern part of Kangwha Island , Korea, was studied. Its caraace breadth, the carapace length , and body height were checked , and the measured raw data were analysed for the growth ratios by the mode subtraction method from the frequency distribution compared with the probability graphs. The growth rate of the crab rose more in summer than in fall and exceeded 10% during the same period on Chojijin tidal flat in Kangwha Island , Korea. Four size groups of the crab appeared in June, 1983 , but they were put together into only on emixed group by autumn. The rapid growth of the crab in summer in middle latitude comes from the seasonal bio-rhythm and the best conditions for food. But the physiological activities of the crab are reduced from autumn and the growth rate reaches the upper limits in many individuals on this tidal flat. The more the crabs grow, the more the breadth ratios to the carapac length become larger and the slopes of correlation curve get smaller. The measured values from the carapace breadth of the crab are more useful for morphological studies than those values from the carapace length or body height.

  • PDF

Mitochondrial OXPHOS genes provides insights into genetics basis of hypoxia adaptation in anchialine cave shrimps

  • Guo, Huayun;Yang, Hao;Tao, Yitao;Tang, Dan;Wu, Qiong;Wang, Zhengfei;Tang, Boping
    • Genes and Genomics
    • /
    • v.40 no.11
    • /
    • pp.1169-1180
    • /
    • 2018
  • Cave shrimps from the genera Typhlatya, Stygiocaris and Typhlopatsa (TST complex) comprises twenty cave-adapted taxa, which mainly occur in the anchialine environment. Anchialine habitats may undergo drastic environmental fluctuations, including spatial and temporal changes in salinity, temperature, and dissolved oxygen content. Previous studies of crustaceans from anchialine caves suggest that they have possessed morphological, behavioral, and physiological adaptations to cope with the extreme conditions, similar to other cave-dwelling crustaceans. However, the genetic basis has not been thoroughly explored in crustaceans from anchialine habitats, which can experience hypoxic regimes. To test whether the TST shrimp-complex hypoxia adaptations matched adaptive evolution of mitochondrial OXPHOS genes. The 13 OXPHOS genes from mitochondrial genomes of 98 shrimps and 1 outgroup were examined. For each of these genes was investigated and compared to orthologous sequences using both gene (i.e. branch-site and Datamonkey) and protein (i.e. TreeSAAP) level approaches. Positive selection was detected in 11 of the 13 candidate genes, and the radical amino acid changes sites scattered throughout the entire TST complex phylogeny. Additionally, a series of parallel/convergent amino acid substitutions were identified in mitochondrial OXPHOS genes of TST complex shrimps, which reflect functional convergence or similar genetic mechanisms of cave adaptation. The extensive occurrence of positive selection is suggestive of their essential role in adaptation to hypoxic anchialine environment, and further implying that TST complex shrimps might have acquired a finely capacity for energy metabolism. These results provided some new insights into the genetic basis of anchialine hypoxia adaptation.