Acknowledgement
Authors thank the anonymous reviewers for their helpful comments on this manuscript. We also thank Juhyun Kim, Sojeong Lee, and Khulan Sharavdorj of Daegu University, and local forestry officers of Lao PDR for their help with field work. We are grateful to the herbarium staffs at BKF, BK, and QBG for their help with herbarium work. This work was supported by a fund from the National Institute of Biological Resources (NIBR), entitled "Survey and Classification of Biological Resources of Mekong Area" (NIBR202006203).
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